In most fish, and in all the rudest types, the swim-bladder is merely a float or balloon, which can be filled with air, and compressed or expanded, so as to make the animal rise or sink at pleasure. But many fish exist in tropical ponds and shallow swamps to whom what has happened artificially to the carp in my friend’s ornamental12 water happens naturally every dry season; the marshy13 sheets in which they live evaporate altogether, and they are therefore compelled to lie dormant14 in the mud without food or drink for many weeks at a time. Under these peculiar15 circumstances, their air-bladder has gradually developed into a true lung; and, what is odder still, we possess in various countries distinct specimens16 at all the intermediate stages from air-bladder to lung in proportion as the ponds which they haunt become dry for longer or shorter periods. The bow-fin of the United States, for example, lives in turbid17 waters which do not quite dry up, but it has acquired the habit of rising to the surface every now and then, and gulping in large mouthfuls of air, which enter its swim-bladder. It does so most frequently when the water is foul18, and there has been little rainfall—in other words, when there is a scarcity19 of oxygen. Accordingly, its air-bladder—though not yet a true lung—is spongy and cellular20 in structure, being adapted for a?rating the blood that passes through it. The mud-fish of Queensland, again, to take a further stage, is a six-foot-long fish which inhabits loaded streams, where its gills do not suffice it for proper respiration21; it has therefore altered its swim-bladder into a rudimentary lung, more advanced than the bow-fin’s, and full of air-cells, richly supplied with blood-vessels, but consisting still of a single cavity. Nevertheless, even this imperfect lung enables the mud-fish to stroll away from its native streams at night, and wander at large on dry land by means of fins22 which are almost legs, and which act like the sprawling23 limbs of certain southern lizards24. In that unnatural25 environment it browses26 on green leaves, and otherwise behaves in a most unfishlike manner. Finally, to complete our rough survey, the African lepidosiren makes its home in waters which dry up completely during the hot season; and it therefore hibernates27 (or rather, ?stivates) for months together in a cocoon28 of hard mud, where it breathes at its ease by means of true lungs, completely divided into lateral29 halves, and approaching in structure those of an air-breathing reptile30.
This interesting series of living evolutionary31 fossils—links that are not missing—is completed for us in some ways by the frogs and toads32, which recapitulate33, as it were, in their own lifetime just such an ancestral developmental history. Each of them begins life as essentially34 a fish—that is to say, as a tadpole35 breathing oxygen dissolved in water, by means of gills, and possessed36 of no limbs for terrestrial locomotion37; he ends it as essentially a full-grown land-reptile, breathing atmospheric38 air by means of lungs, having discarded his now needless fin-fringed tail, and possessed of jumping legs of great muscular power. And the metamorphosis he thus undergoes answers exactly to just such a drying-up of the ponds that bore him. In early spring, when the temporary puddles39 are full of water, the parent frogs lay their spawn40 by hundreds in the ancestral element; and soon the little black tadpoles—true fish of a primitive41 type in all but name—swarm forth42 and swim in seething43 masses in the momentary44 medium. But as the sun begins to dry up the water in their dwelling-place they lose their fins and gills, pass from fish to amphibians45, and shortly hop46 ashore47, provided with four legs and a pair of lungs specially48 adapted for direct air-breathing. There we have a marvellous piece of evolutionary magic still going on every day before our eyes, which would sound incredible to us if a man of science reported it for the first time from Central Africa or New Guinea. The frog, in short, shows us successively in his own person the self-same stages of development which the various mud-fish preserve for us in distant regions, as types of distinct and unrelated species.

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1
puddle
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n.(雨)水坑,泥潭 | |
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2
inadequate
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adj.(for,to)不充足的,不适当的 | |
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3
lustreless
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adj.无光泽的,无光彩的,平淡乏味的 | |
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4
suffocating
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a.使人窒息的 | |
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5
gasping
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adj. 气喘的, 痉挛的 动词gasp的现在分词 | |
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6
gulping
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v.狼吞虎咽地吃,吞咽( gulp的现在分词 );大口地吸(气);哽住 | |
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7
misery
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n.痛苦,苦恼,苦难;悲惨的境遇,贫苦 | |
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8
instinctive
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adj.(出于)本能的;直觉的;(出于)天性的 | |
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9
nay
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adv.不;n.反对票,投反对票者 | |
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10
mechanism
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n.机械装置;机构,结构 | |
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11
inhaling
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v.吸入( inhale的现在分词 ) | |
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12
ornamental
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adj.装饰的;作装饰用的;n.装饰品;观赏植物 | |
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13
marshy
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adj.沼泽的 | |
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14
dormant
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adj.暂停活动的;休眠的;潜伏的 | |
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15
peculiar
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adj.古怪的,异常的;特殊的,特有的 | |
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16
specimens
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n.样品( specimen的名词复数 );范例;(化验的)抽样;某种类型的人 | |
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17
turbid
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adj.混浊的,泥水的,浓的 | |
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18
foul
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adj.污秽的;邪恶的;v.弄脏;妨害;犯规;n.犯规 | |
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19
scarcity
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n.缺乏,不足,萧条 | |
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20
cellular
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adj.移动的;细胞的,由细胞组成的 | |
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21
respiration
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n.呼吸作用;一次呼吸;植物光合作用 | |
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22
fins
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[医]散热片;鱼鳍;飞边;鸭掌 | |
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23
sprawling
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adj.蔓生的,不规则地伸展的v.伸开四肢坐[躺]( sprawl的现在分词 );蔓延;杂乱无序地拓展;四肢伸展坐着(或躺着) | |
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24
lizards
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n.蜥蜴( lizard的名词复数 ) | |
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25
unnatural
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adj.不自然的;反常的 | |
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26
browses
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n.吃草( browse的名词复数 );随意翻阅;(在商店里)随便看看;(在计算机上)浏览信息v.吃草( browse的第三人称单数 );随意翻阅;(在商店里)随便看看;(在计算机上)浏览信息 | |
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27
hibernates
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(某些动物)冬眠,蛰伏( hibernate的第三人称单数 ) | |
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28
cocoon
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n.茧 | |
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29
lateral
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adj.侧面的,旁边的 | |
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30
reptile
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n.爬行动物;两栖动物 | |
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31
evolutionary
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adj.进化的;演化的,演变的;[生]进化论的 | |
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32
toads
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n.蟾蜍,癞蛤蟆( toad的名词复数 ) | |
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33
recapitulate
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v.节述要旨,择要说明 | |
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34
essentially
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adv.本质上,实质上,基本上 | |
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35
tadpole
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n.[动]蝌蚪 | |
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36
possessed
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adj.疯狂的;拥有的,占有的 | |
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37
locomotion
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n.运动,移动 | |
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38
atmospheric
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adj.大气的,空气的;大气层的;大气所引起的 | |
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39
puddles
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n.水坑, (尤指道路上的)雨水坑( puddle的名词复数 ) | |
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40
spawn
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n.卵,产物,后代,结果;vt.产卵,种菌丝于,产生,造成;vi.产卵,大量生产 | |
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41
primitive
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adj.原始的;简单的;n.原(始)人,原始事物 | |
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42
forth
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adv.向前;向外,往外 | |
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43
seething
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沸腾的,火热的 | |
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44
momentary
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adj.片刻的,瞬息的;短暂的 | |
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45
amphibians
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两栖动物( amphibian的名词复数 ); 水陆两用车; 水旱两生植物; 水陆两用飞行器 | |
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46
hop
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n.单脚跳,跳跃;vi.单脚跳,跳跃;着手做某事;vt.跳跃,跃过 | |
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47
ashore
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adv.在(向)岸上,上岸 | |
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48
specially
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adv.特定地;特殊地;明确地 | |
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