Here, on the edge of the down, where I have come up to get a good blowing from the clear south-west breeze, I have just sat down to rest myself awhile and to admire the view, and have reverted4 for a moment to my old habit of snail5-hunting. Years ago, when evolution was an infant—an infant much troubled by the complaints inseparable from infancy6, but still a sturdy and vigorous child, destined7 to outlive and outgrow8 its early attacks—I used to collect slugs and snails9, from an evolutionist standpoint, and put their remains10 into a cabinet; and to this day I seldom go out for a walk without a few pill-boxes in my pocket, in case I should happen to hit upon any remarkable11 specimen12. Now here in the tall moss13 which straggles over an old heap of stones I have this moment lighted upon a beautifully marked shell of our prettiest English snail. How beautiful it is I could hardly make you believe, unless I had you here and could show it to you; for most people only know the two or three ugly brown or banded snails that prey14 upon their cabbages and lettuces15, and have no notion of the lovely shells to be found by hunting among English copses and under the dead leaves of Scotch16 hill-sides. This cyclostoma, however,—I must trouble you with a Latin name for once—is so remarkably17 pretty, with its graceful18 elongated19 spiral whorls, and its delicately chiselled20 fretwork tracery, that even naturalists21 (who have perhaps, on the whole, less sense of beauty than any class of men I know) have recognised its loveliness by giving it the specific epithet22 of elegans. It is big enough for anybody to notice it, being about the size of a periwinkle; and its exquisite23 stippled24 chasing is strongly marked enough to be perfectly25 visible to the naked eye. But besides its beauty, the cyclostoma has a strong claim upon our attention because of its curious history.
Long ago, in the infantile days of evolutionism, I often wondered why people made collections on such an irrational26 plan. They always try to get what they call the most typical specimens27, and reject all those which are doubtful or intermediate. Hence the dogma of the fixity of species becomes all the more firmly settled in their minds, because they never attend to the existing links which still so largely bridge over the artificial gaps created by our nomenclature between kind and kind. I went to work on the opposite plan, collecting all those aberrant28 individuals which most diverged29 from the specific type. In this way I managed to make some series so continuous that one might pass over specimens of three or four different kinds, arranged in rows, without ever being able to say quite clearly, by the eye alone, where one group ended and the next group began. Among the snails such an arrangement is peculiarly easy; for some of the species are very indefinite, and the varieties are numerous under each species. Nothing can give one so good a notion of the plasticity of organic forms as such a method. The endless varieties and intermediate links which exist amongst dogs is the nearest example to it with which ordinary observers are familiar.
But the cyclostoma is a snail which introduces one to still deeper questions. It belongs in all our scientific classifications to the group of lung-breathing mollusks, like the common garden snail. Yet it has one remarkable peculiarity30: it possesses an operculum, or door to its shell, like that of the periwinkle. This operculum represents among the univalves the under-shell of the oyster31 or other bivalves; but it has completely disappeared in most land and fresh-water snails, as well as among many marine32 species. The fact of its occurrence in the cyclostoma would thus be quite inexplicable33 if we were compelled to regard it as a descendant of the other lung-breathing mollusks. So far as I know, all naturalists have till lately always so regarded it; but there can be very little doubt, with the new light cast upon the question by Darwinism, that they are wrong. There exists in all our ponds and rivers another snail, not breathing by means of lungs, but provided with gills, known as paludina. This paludina has a door to its shell, like the cyclostoma; and so, indeed, have all its allies. Now, strange as it sounds to say so, it is pretty certain that we must really class this lung-breathing cyclostoma among the gill-breathers, because of its close resemblance to the paludina. It is, in fact, one of these gill-breathing pond-snails which has taken to living on dry land, and so has acquired the habit of producing lungs. All molluscan lungs are very simple: they consist merely of a small sac or hollow behind the head, lined with blood-vessels; and every now and then the snail opens this sac, allowing the air to get in and out by natural change, exactly as when we air a room by opening the windows. So primitive35 a mechanism36 as this could be easily acquired by any soft-bodied animal like a snail. Besides, we have many intermediate links between the pond-snails and my cyclostoma here. There are some species which live in moist moss, or the beds of trickling37 streams. There are others which go further from the water, and spend their days in damp grass. And there are yet others which have taken to a wholly terrestrial existence in woods or meadows and under heaps of stones. All of them agree with the pond-snails in having an operculum, and so differ from the ordinary land and river snails, the mouths of whose shells are quite unprotected. Thus land-nails have two separate origins—one large group (including the garden-snail) being derived38 from the common fresh-water mollusks, while another much smaller group (including the cyclostoma) is derived from the operculated pond-snails.
How is it, then, that naturalists had so long overlooked this distinction? Simply because their artificial classification is based entirely39 upon the nature of the breathing apparatus40. But, as Mr. Wallace has well pointed41 out, obvious and important functional42 differences are of far less value in tracing relationship than insignificant43 and unimportant structural44 details. Any water-snail may have to take to a terrestrial life if the ponds in which it lives are liable to dry up during warm weather. Those individuals alone will then survive which display a tendency to oxygenise their blood by some rudimentary form of lung. Hence the possession of lungs is not the mark of a real genealogical class, but a mere34 necessary result of a terrestrial existence. On the other hand, the possession of an operculum, unimportant as it may be to the life of the animal, is a good test of relationship by descent. All snails which take to living on land, whatever their original form, will acquire lungs: but an operculated snail will retain its operculum, and so bear witness to its ancestry45; while a snail which is not operculated will of course show no tendency to develop such a structure, and so will equally give a true testimony46 as to its origin. In short, the less functionally47 useful any organ is, the higher is its value as a gauge48 of its owner's pedigree, like a Bourbon nose or an Austrian lip.
点击收听单词发音
1 standing | |
n.持续,地位;adj.永久的,不动的,直立的,不流动的 | |
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2 miserable | |
adj.悲惨的,痛苦的;可怜的,糟糕的 | |
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3 foliage | |
n.叶子,树叶,簇叶 | |
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4 reverted | |
恢复( revert的过去式和过去分词 ); 重提; 回到…上; 归还 | |
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5 snail | |
n.蜗牛 | |
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6 infancy | |
n.婴儿期;幼年期;初期 | |
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7 destined | |
adj.命中注定的;(for)以…为目的地的 | |
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8 outgrow | |
vt.长大得使…不再适用;成长得不再要 | |
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9 snails | |
n.蜗牛;迟钝的人;蜗牛( snail的名词复数 ) | |
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10 remains | |
n.剩余物,残留物;遗体,遗迹 | |
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11 remarkable | |
adj.显著的,异常的,非凡的,值得注意的 | |
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12 specimen | |
n.样本,标本 | |
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13 moss | |
n.苔,藓,地衣 | |
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14 prey | |
n.被掠食者,牺牲者,掠食;v.捕食,掠夺,折磨 | |
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15 lettuces | |
n.莴苣,生菜( lettuce的名词复数 );生菜叶 | |
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16 scotch | |
n.伤口,刻痕;苏格兰威士忌酒;v.粉碎,消灭,阻止;adj.苏格兰(人)的 | |
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17 remarkably | |
ad.不同寻常地,相当地 | |
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18 graceful | |
adj.优美的,优雅的;得体的 | |
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19 elongated | |
v.延长,加长( elongate的过去式和过去分词 ) | |
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20 chiselled | |
adj.凿过的,凿光的; (文章等)精心雕琢的v.凿,雕,镌( chisel的过去式 ) | |
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21 naturalists | |
n.博物学家( naturalist的名词复数 );(文学艺术的)自然主义者 | |
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22 epithet | |
n.(用于褒贬人物等的)表述形容词,修饰语 | |
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23 exquisite | |
adj.精美的;敏锐的;剧烈的,感觉强烈的 | |
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24 stippled | |
v.加点、绘斑,加粒( stipple的过去式和过去分词 );(把油漆、水泥等的表面)弄粗糙 | |
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25 perfectly | |
adv.完美地,无可非议地,彻底地 | |
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26 irrational | |
adj.无理性的,失去理性的 | |
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27 specimens | |
n.样品( specimen的名词复数 );范例;(化验的)抽样;某种类型的人 | |
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28 aberrant | |
adj.畸变的,异常的,脱离常轨的 | |
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29 diverged | |
分开( diverge的过去式和过去分词 ); 偏离; 分歧; 分道扬镳 | |
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30 peculiarity | |
n.独特性,特色;特殊的东西;怪癖 | |
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31 oyster | |
n.牡蛎;沉默寡言的人 | |
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32 marine | |
adj.海的;海生的;航海的;海事的;n.水兵 | |
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33 inexplicable | |
adj.无法解释的,难理解的 | |
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34 mere | |
adj.纯粹的;仅仅,只不过 | |
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35 primitive | |
adj.原始的;简单的;n.原(始)人,原始事物 | |
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36 mechanism | |
n.机械装置;机构,结构 | |
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37 trickling | |
n.油画底色含油太多而成泡沫状突起v.滴( trickle的现在分词 );淌;使)慢慢走;缓慢移动 | |
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38 derived | |
vi.起源;由来;衍生;导出v.得到( derive的过去式和过去分词 );(从…中)得到获得;源于;(从…中)提取 | |
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39 entirely | |
ad.全部地,完整地;完全地,彻底地 | |
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40 apparatus | |
n.装置,器械;器具,设备 | |
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41 pointed | |
adj.尖的,直截了当的 | |
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42 functional | |
adj.为实用而设计的,具备功能的,起作用的 | |
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43 insignificant | |
adj.无关紧要的,可忽略的,无意义的 | |
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44 structural | |
adj.构造的,组织的,建筑(用)的 | |
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45 ancestry | |
n.祖先,家世 | |
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46 testimony | |
n.证词;见证,证明 | |
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47 functionally | |
adv.机能上地,官能地 | |
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48 gauge | |
v.精确计量;估计;n.标准度量;计量器 | |
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