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CHAPTER II. THE FUNCTIONAL RELATIONS OF THE NERVOUS SYSTEM.
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 The distinguishable parts of this system are the central axis1, the cranial nerves, and the spinal2 nerves, with the chain of ganglia and nerves composing the Sympathetic. Let us briefly3 set down what is known of their special offices.
Men very early discovered that the nerves were in some way ministrant to Sensation and Movement; a divided nerve always being accompanied by insensibility and immobility in the limb. Galen, observing that paralysis4 of movement sometimes occurred without insensibility, suggested that there were two kinds of nerve; but no one177 was able to furnish satisfactory evidence in support of this suggestion until early in the present century, when the experiments of Charles Bell, perfected by those of Majendie and Müller, placed the suggestion beyond dispute.
 
Fig5. 12.—Transverse sections of spinal cord (dorsal6 region).
24. Fig. 12 is a diagram (not a drawing of the actual aspect, which would be hardly intelligible7 to readers unversed in such matters) representing two transverse sections of the spinal cord just where the nerve-roots issue. The gray substance is somewhat in the form of a rude H, in the dorsal region, and of the expanded wings of a butterfly in the lumbar enlargements (Figs. 4–6); the extremities8 of this gray substance are the anterior9 and posterior horns. We have already said that from the anterior horns of each half issue the roots of the motor nerves, which pass to the muscles. From the posterior horns issue the sensory10 nerves, which, soon after leaving the cord, enter the ganglia before joining the motor nerves, and then pass to the skin, in the same sheath with their companions, separating again as they reach the muscles and surfaces where they are to be distributed. When this mixed nerve is cut through, or tied, all sensation and movement disappear from the parts innervated. But if only one of the roots be cut through, above the ganglion, there will then be only a loss of movement or a loss of sensation. Thus suppose the section be made at a, b, A: we have then divided a sensory nerve, and no pinching or pricking11 of the part innervated by that nerve will be felt; but movement will take place if the under nerve be irritated, or if a sensation elsewhere be excited. Now reverse the experiment, as at B, c, d. Then, pricking of the skin will be felt, but no movement will respond. The nerve which enters the cord at the upper (posterior) part is therefore a sensory nerve; that which enters at the under (anterior) part is motor. The direction is in each case indicated by178 the arrow. The central end b, if irritated, will produce sensation; whereas the peripheral12 end a produces neither sensation nor movement. The central end d produces neither sensation nor movement; the peripheral end c produces movement.
25. Two facts are proved by these experiments. First, that the co-operation of the centre is necessary for Sensation, but not for Movement. Although normally all the muscles of the trunk are moved only when their centre has been excited, yet any irritation13 applied14 directly to the muscle nerve, even when separated from its centre, produces a movement. And to this we may add that a slighter stimulus15 will move the muscle by direct irritation of the nerve, than by indirect irritation through the centre; a slighter stimulus also will suffice when applied to the nerve than when applied to the muscle itself.
26. The second fact proved is known as Bell’s Law, that the sensory and motor channels are respectively the posterior and anterior nerves. The fact is indisputable, but its theoretic interpretation16 can no longer be accepted in its original form. Bell supposed the two nerves to be different in kind, endowed with different specific energies, the one sensitive, the other motor. The majority of writers still express themselves as if they adopted this view. We shall, however, presently see reason for replacing it by the more consistent interpretation which assigns one and the same property to both nerves, marking their distinction by the terms afferent and efferent; the one set being anatomically so disposed that it conveys stimuli17 from the surfaces to the centre, and the other set conveying stimuli from the centre to the muscles, glands18, and other cells.89
179 27. Bell’s discovery was rapidly generalized. The principle of localization was extended to all nerves, and of course to the posterior and anterior columns of the spinal cord, which indeed were assumed to be continuations of the nerves. Bell, who was greater as an anatomist than as a philosopher, always maintained that anatomical deduction19 was superior to experiment. But this was to misunderstand the reach of deduction, which is only valid20 to the extent of its premises21.90 In the present case, the premises assumed that the posterior columns were continuations of the posterior roots, and carried impressions to the brain, the anterior columns carrying back from the brain the “mandates of the will.” Experiment has, however, decisively shown that it is not through the posterior columns that sensory impressions travel to the brain, but through the central gray substance.
28. The spinal cord with its central gray substance is at each point a centre of reflexion. Connected as it is with different organs, we artificially consider it as a chain of different centres, and try to detect the functional22 relations of its parts. The inquiry23 is important, but we must bear in mind the cardinal24 principle that diversity of Function depends on the organs innervated, and not on a diversity of Property in the nervous tissue. Although all nerves have a common structure and common property, yet we distinguish them as sensory and motor; and the sensory we subdivide25 into those of Special Sensation and those of Systemic Sensation. The motor we divide into muscular, vasomotor, and glandular26. The hypothesis of specific energies must be relinquished27 (§ 63).
180 In like manner all centres have a common structure and a common property, with a great diversity of functional relations. Here also the hypothesis of specific energies has been generally adopted, owing to a mistaken conception of the biological principle just mentioned. The cerebral28 hemispheres are credited with the properties of sensation, thought, and volition29; the cerebellum with the property of muscular co-ordination; the spinal cord with the property of reflexion.
29. No attempt to assign the true functional relations of the centres will be made at the present stage of our exposition. We must learn more of the processes in Sensation, Thought, and Volition, before we can unravel30 the complex physiological31 web on which they depend. But here, provisionally, may be set down what observation and experiment have disclosed respecting the part played by certain centres. We know, for example, that when the cerebral hemispheres are carefully removed from a reptile32 or a bird, all the essentially33 vital functions go on pretty much as before, but a great disturbance34 in some of the psychical35 functions is observed. The brainless bird eats, drinks, sleeps, moves its limbs separately and in combination, manifests sensibility to light, sound, and touch, performs such instinctive36 actions as preening37 its feathers, or thrusting the head under the wing while roosting. Throw it into the air and it will fly. In its flight it will avoid obstacles, and will alight upon a ledge38, or your shoulder. But it will not fly unless thrown into the air; it will not escape through the open door or window; it will avoid objects, but will show no fear of them,—alighting on your head, for example, without hesitation39. It is sensitive to light, and may in a certain sense be said to see; but it fails to perceive what is seen. It will eat and drink, if food and water be administered, but it will starve near a heap of grain and never peck it, not181 even if the beak40 be thrust into the heap. A grain, or strip of meat, may be thrust inside the beak; there it will remain unswallowed, unless it touches the back of the mouth, then swallowing at once follows the stimulus. The bird with its brain will fly away if you turn the finger, or stick, on which it is perching; without its brain, it makes no attempt to fly, but flutters its wings, and balances itself. If you open the mouth of a cat, or rabbit, and drop in some bitter fluid, the animal closes its mouth firmly, and resists your efforts to repeat the act; without its brain, the animal shows the same disgust at the taste, but never resists the preliminaries of the repetition.
30. These, and analogous41 facts, have been noted42 by various experimenters. They are very far from proving what is usually concluded; but they prove the important negative position that the cerebrum is not the centre of innervation for any of the organs on which the observed actions depend. Thus, the cerebrum is not necessary to sight: ergo it does not innervate the eye. It is not necessary to hearing: ergo it does not innervate the ear.91 It is not necessary to breathing, swallowing, flying, etc.: ergo it does not innervate the organs of these functions.
What then is lost? We have only to remember that the cerebrum is continuous with the thalami and corpora striata, and, through its crura, with the medulla oblongata and medulla spinalis, to foresee that its removal must more or less affect the whole neural43 axis, and consequently disturb the actions of the whole organism; this disturbance will often have the appearances which would182 be due to the removal of a central apparatus44, so that we shall be apt to attribute the cessation of a function to the loss of its organ, when in fact the cessation is due simply to an arrest of the organ by irritation. Thus the cessation of consciousness, or of any particular movements, when the cerebrum is removed, is no decisive proof that the cerebrum is the organ of consciousness, or of the movement in question. This point will be duly considered hereafter. What we have now to consider is the facts observed after removal of the cerebrum.
First, we observe a loss of that power of combining present states with past states, present feelings with feelings formerly45 excited in conjunction with them, the power which enables the animal to adjust its actions to certain sensations now unfelt but which will be felt in consequence of the adjustment. Secondly46, we observe a loss of Spontaneity: the bird, naturally mobile and alert, now sits moveless for hours in a sort of stupor47, occasionally preening its feathers, but rarely quitting its resting-place. All the most conspicuous48 phenomena49 which we assign to Intelligence and Will seem absent. The sensations are altered and diminished. Many Instincts have disappeared, but some remain. The sexual feeling is preserved, although the bird has lost all power of directing its actions so as to gratify the desire. But these effects are only observed when the whole of both hemispheres have been removed. If a small portion remain the bird retains most of its faculties50, though with less energy. In frogs and fishes there is little discernible effect observed when a large portion of the cerebrum is removed.
31. Now take away from this mutilated bird its cerebellum: all the functions continue as before except that some combined movements can no longer be effected; flight is impossible; walking is a mere51 stagger. Remove183 only the lateral52 lobes53, and though flight is still possible great incoherence of the wings is observed, whereas walking is not much affected54. If only the cerebellum be removed, the cerebrum being intact, the phenomena are very different. All the perceptions and almost all the emotions, all the spontaneity and vivacity55 are retained; but the sexual instinct, which was manifested when the cerebrum was removed, is now quite gone. What we call Intelligence seems unaffected. The bird hears, and understands the meaning of the sounds, sees and perceives, sees and fears, sees and adjusts its movements with a mental vision of unseen consequences.92
32. Are we from these facts to conclude that the cerebrum is the “organ of the mind”; that it is “the seat” of sensation, thought, emotion, volition; and that the cerebellum is the “seat” of the sexual instinct, and muscular co-ordination? Such conclusions have found acceptance, even from physiologists56 who would have been startled had any one ventured to affirm that the medulla oblongata was the “organ” of Respiration57, because Respiration ceases when this centre is destroyed. I shall have to combat this notion at various stages of my exposition. Here let me simply say that it is irreconcilable58 with any clear conception of organ and function; and is plainly irreconcilable with any survey of psychical phenomena in animals in whom the cerebrum does not exist, and in animals from whom it has been removed.
What the facts indisputably prove is that the cerebrum has an important part in the mechanism59 by which the most complex psychical combinations are effected, and that the cerebellum has an important part in the mechanism by which the most complex muscular combinations184 are effected. The supreme60 importance of the cerebrum may be inferred from its dominating all the other centres, and from its preponderance in size. In man it stands to all the other cranial centres together in the relation of 11 to 3. It is about five times as heavy as the spinal cord—that is to say from 1,100 to 1,400 grammes, compared with 27 to 30 grammes. The quantity of blood circulating through it is immense. Haller estimated the cranial circulation as one fifth of the whole circulation. If, therefore, the Nervous Centres are agents in the production of Sensation and Intelligence, by far the largest share must be allotted61 to the cranial centres, and of these the largest to the Cerebrum.
33. It is, however, one thing to recognize the Cerebrum as having an important part in the production of psychical phenomena, another thing to localize all the phenomena in it as their organ and seat—a localization which soon becomes even more absurd, when of all the cerebral structure the multipolar cells alone are admitted as the active agents!
As was said just now, we recognize in the Medulla Oblongata the nervous centre of Respiration, but we do not suppose that Respiration has its seat there, nor that this centre is absolutely indispensable for the essential part of the process. We respire by our skin, as well as by our lungs; many animals respire who have nothing like a medulla oblongata; as many animals feel, and manifest will, who have nothing like a cerebrum. The destruction of centres is of course a disturbance of the mechanisms62 which they regulate. But even the observed results of a destruction require very close examination, and are liable to erroneous interpretations63. The disappearance64 of a function following the destruction, or disease of a particular part, is not to be accepted as a proof that this part is the organ of the lost function; because precisely185 the same phenomena may often be observed following the destruction of a totally different part.93 But one result may always be relied on, and that is the persistence65 of a function after removal of a particular part. Thus there is a certain spot of the cerebral convolutions from which movements of the limbs are excited when the electrodes are applied to it; removal of the substance is immediately followed by paralysis of the limbs. Are we to conclude that this spot is the organ of the function? It is true that the function is called into action by a stimulus applied to this spot: true that the function suddenly vanishes when the substance of this spot is destroyed. Nevertheless, what seems a loss of function is only a disturbance. In two or three days the paralysis begins to disappear, and at the end of a week the limbs are moved nearly in the normal manner. And the same is true when the spot in question is destroyed on both sides. The recovery of the function shows that the absent part was not its organ. There is a paradoxical experiment recorded by M. Paul Bert which may be cited here. He removed the right cerebral hemisphere from a chameleon66, and found that the limbs on the left side were paralyzed; but on his then removing the left cerebral hemisphere the limbs of the left side recovered their activity. A similar result was obtained by Lussana and Lemoigne by extirpation67 of the thalami. When we find combined movements persisting after the cerebellum has been destroyed, we may be sure that the cerebellum is186 not the organ by which such combinations take place; and when we find sensation and volition manifested after the cerebrum has been removed, we may be sure that the cerebrum is not the organ for these sensations and volitions.
34. And this we do find. Physiologists, indeed, for the most part, deny it; or rather, while they admit the observed facts, they refuse to admit the only consistent interpretation, biassed68 as they are by the traditional conception of the brain. After having for many years persistently69 denied Sensibility to any centre except the cerebrum, they are now generally agreed in including the medulla oblongata within the privileged region; but they still exclude the medulla spinalis.
35. If all the cranial centres as far as the medulla oblongata are removed from young rabbits, dogs, or cats, there are unmistakable evidences of Sensibility in their cries when their tails are pinched, their moving jaws70 (as in mastication) when bitters are placed in their mouths, and their raised paws rubbing their noses, when irritating vapors71 are applied. It is said indeed that the cries are no signs of pain; and this is probable; but they are assuredly signs of Sensibility.
35. The frog thus mutilated has lost indeed all its special senses, except Touch, but it still breathes, struggles when grasped, thrusts aside the pincers which irritate it, or wipes away acid dropped on its skin. If the eye be lightly touched, the eyelid72 closes; if the touch be repeated three or four times, the foreleg is raised to push the irritant away; if still repeated, the head is turned aside; but however prolonged the irritation, the frog neither hops73, nor crawls away, as he does when the cerebellum remains74. Place the brainless frog on his back, and if the medulla oblongata remains he will at once regain75 the normal position; but if that part is absent he will lie helpless on his back. The power of preserving equilibrium76 in difficult187 positions—which of course implies a nice co-ordination of muscles—resides in the so-called optic lobes of the frog (what in mammals are called the corpora quadrigemina).
37. With the destruction of each part of the central mass there will necessarily be some disturbance of the mechanism; but difficult as may be the task of detecting by experiment what is the normal action of any one part, there ought to be no hesitation in recognizing the persistence of functions after certain parts are destroyed. The spinal cord is anatomically known to be the centre from which the limbs, trunk, and genito-urinary organs are innervated. So long as the mechanism of the actions involving such organs is intact, no removal of other parts will prevent this mechanism from exhibiting its normal action. There may indeed arise, and there has arisen, the doubt whether Sensibility is involved in the action of any nerve centre below the medulla oblongata. But this doubt is founded on the traditional hypothesis respecting the seat of Sensation, and is flagrantly at variance77 with the logical conclusions of Anatomy78 and Experiment.
38. Anatomy shows that the structure of the spinal cord is in all essential characters the same as that of the medulla oblongata; and indeed that the whole central axis has one continuous tissue, somewhat variously arranged, and in relation with various organs.
Abundant Experiment has shown that the spinal cord, apart from the encephalon, is capable of acting79 as a sensorial and volitional80 centre. The striking facts advanced by Pflüger, Auerbach, and myself, have not been impugned;94 but their interpretation has been generally188 rejected. We showed that a brainless frog responded to stimulation81 in actions which bore so close a resemblance to actions admitted to be sensorial and volitional—showed the frog adapting itself to new conditions, and acquiring dexterity82 in executing actions which at first were impossible or difficult, devising combinations to effect a purpose which never by any possibility could have formed part of its habits—manifesting, in a word, such signs of Sensibility, that no one witnessing the experiments could hesitate as to the interpretation, had he not been biassed by the traditions of the schools.
39. Our opponents argued that in spite of all appearances there were profound differences between the actions of the normal and the brainless animal, and that the latter were due simply to Reflex Action. I also insist on profound differences; but underlying83 these there are fundamental identities. As to the Reflex Action, two points will hereafter be brought forward: 1°, that all central action is reflex, the cerebral no less than the spinal; 2°, that the hypothesis of Reflex Action being purely84 mechanical, and distinguished85 from Voluntary Action in not involving Sensibility, is an hypothesis which must be relinquished.
40. Postponing86, however, all discussion of these points, let me here say that the doctrine87 maintained in these pages is that the whole cerebro-spinal axis is a centre of Reflexion, its various segments taking part in the performance of different kinds of combined action. It has one common property, Sensibility; and different parts of it minister to different functions—the optic centre being different from the auditory, the cerebral from the spinal; and so on. To make this intelligible, however, we must first learn what is known respecting the properties of nerve-tissue.

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1 axis sdXyz     
n.轴,轴线,中心线;坐标轴,基准线
参考例句:
  • The earth's axis is the line between the North and South Poles.地轴是南北极之间的线。
  • The axis of a circle is its diameter.圆的轴线是其直径。
2 spinal KFczS     
adj.针的,尖刺的,尖刺状突起的;adj.脊骨的,脊髓的
参考例句:
  • After three days in Japan,the spinal column becomes extraordinarily flexible.在日本三天,就已经使脊椎骨变得富有弹性了。
  • Your spinal column is made up of 24 movable vertebrae.你的脊柱由24个活动的脊椎骨构成。
3 briefly 9Styo     
adv.简单地,简短地
参考例句:
  • I want to touch briefly on another aspect of the problem.我想简单地谈一下这个问题的另一方面。
  • He was kidnapped and briefly detained by a terrorist group.他被一个恐怖组织绑架并短暂拘禁。
4 paralysis pKMxY     
n.麻痹(症);瘫痪(症)
参考例句:
  • The paralysis affects his right leg and he can only walk with difficulty.他右腿瘫痪步履维艰。
  • The paralysis affects his right leg and he can only walk with difficulty.他右腿瘫痪步履维艰。
5 fig L74yI     
n.无花果(树)
参考例句:
  • The doctor finished the fig he had been eating and selected another.这位医生吃完了嘴里的无花果,又挑了一个。
  • You can't find a person who doesn't know fig in the United States.你找不到任何一个在美国的人不知道无花果的。
6 dorsal rmEyC     
adj.背部的,背脊的
参考例句:
  • His dorsal fin was down and his huge pectorals were spread wide.它的脊鳍朝下耷拉着,巨大的胸鳍大张着。
  • The shark's dorsal fin was cut off by the fisherman.鲨鱼的背鳍被渔夫割了下来。
7 intelligible rbBzT     
adj.可理解的,明白易懂的,清楚的
参考例句:
  • This report would be intelligible only to an expert in computing.只有计算机运算专家才能看懂这份报告。
  • His argument was barely intelligible.他的论点不易理解。
8 extremities AtOzAr     
n.端点( extremity的名词复数 );尽头;手和足;极窘迫的境地
参考例句:
  • She was most noticeable, I thought, in respect of her extremities. 我觉得她那副穷极可怜的样子实在太惹人注目。 来自辞典例句
  • Winters may be quite cool at the northwestern extremities. 西北边区的冬天也可能会相当凉。 来自辞典例句
9 anterior mecyi     
adj.较早的;在前的
参考例句:
  • We've already finished the work anterior to the schedule.我们已经提前完成了工作。
  • The anterior part of a fish contains the head and gills.鱼的前部包括头和鳃。
10 sensory Azlwe     
adj.知觉的,感觉的,知觉器官的
参考例句:
  • Human powers of sensory discrimination are limited.人类感官分辨能力有限。
  • The sensory system may undergo long-term adaptation in alien environments.感觉系统对陌生的环境可能经过长时期才能适应。
11 pricking b0668ae926d80960b702acc7a89c84d6     
刺,刺痕,刺痛感
参考例句:
  • She felt a pricking on her scalp. 她感到头皮上被扎了一下。
  • Intercostal neuralgia causes paroxysmal burning pain or pricking pain. 肋间神经痛呈阵发性的灼痛或刺痛。
12 peripheral t3Oz5     
adj.周边的,外围的
参考例句:
  • We dealt with the peripheral aspects of a cost reduction program.我们谈到了降低成本计划的一些外围问题。
  • The hotel provides the clerk the service and the peripheral traveling consultation.旅舍提供票务服务和周边旅游咨询。
13 irritation la9zf     
n.激怒,恼怒,生气
参考例句:
  • He could not hide his irritation that he had not been invited.他无法掩饰因未被邀请而生的气恼。
  • Barbicane said nothing,but his silence covered serious irritation.巴比康什么也不说,但是他的沉默里潜伏着阴郁的怒火。
14 applied Tz2zXA     
adj.应用的;v.应用,适用
参考例句:
  • She plans to take a course in applied linguistics.她打算学习应用语言学课程。
  • This cream is best applied to the face at night.这种乳霜最好晚上擦脸用。
15 stimulus 3huyO     
n.刺激,刺激物,促进因素,引起兴奋的事物
参考例句:
  • Regard each failure as a stimulus to further efforts.把每次失利看成对进一步努力的激励。
  • Light is a stimulus to growth in plants.光是促进植物生长的一个因素。
16 interpretation P5jxQ     
n.解释,说明,描述;艺术处理
参考例句:
  • His statement admits of one interpretation only.他的话只有一种解释。
  • Analysis and interpretation is a very personal thing.分析与说明是个很主观的事情。
17 stimuli luBwM     
n.刺激(物)
参考例句:
  • It is necessary to curtail or alter normally coexisting stimuli.必需消除或改变正常时并存的刺激。
  • My sweat glands also respond to emotional stimuli.我的汗腺对情绪刺激也能产生反应。
18 glands 82573e247a54d4ca7619fbc1a5141d80     
n.腺( gland的名词复数 )
参考例句:
  • a snake's poison glands 蛇的毒腺
  • the sebaceous glands in the skin 皮脂腺
19 deduction 0xJx7     
n.减除,扣除,减除额;推论,推理,演绎
参考例句:
  • No deduction in pay is made for absence due to illness.因病请假不扣工资。
  • His deduction led him to the correct conclusion.他的推断使他得出正确的结论。
20 valid eiCwm     
adj.有确实根据的;有效的;正当的,合法的
参考例句:
  • His claim to own the house is valid.他主张对此屋的所有权有效。
  • Do you have valid reasons for your absence?你的缺席有正当理由吗?
21 premises 6l1zWN     
n.建筑物,房屋
参考例句:
  • According to the rules,no alcohol can be consumed on the premises.按照规定,场内不准饮酒。
  • All repairs are done on the premises and not put out.全部修缮都在家里进行,不用送到外面去做。
22 functional 5hMxa     
adj.为实用而设计的,具备功能的,起作用的
参考例句:
  • The telephone was out of order,but is functional now.电话刚才坏了,但现在可以用了。
  • The furniture is not fancy,just functional.这些家具不是摆着好看的,只是为了实用。
23 inquiry nbgzF     
n.打听,询问,调查,查问
参考例句:
  • Many parents have been pressing for an inquiry into the problem.许多家长迫切要求调查这个问题。
  • The field of inquiry has narrowed down to five persons.调查的范围已经缩小到只剩5个人了。
24 cardinal Xcgy5     
n.(天主教的)红衣主教;adj.首要的,基本的
参考例句:
  • This is a matter of cardinal significance.这是非常重要的事。
  • The Cardinal coloured with vexation. 红衣主教感到恼火,脸涨得通红。
25 subdivide DtGwN     
vt.细分(细区分,再划分,重分,叠分,分小类)
参考例句:
  • You can use sales organizations to subdivide markets into regions.用销售组织将市场细分为区域。
  • The verbs were subdivided into transitive and intransitive categories.动词可细分为及物动词和不及物动词。
26 glandular wgExR     
adj.腺体的
参考例句:
  • Terry has been laid low with glandular fever for nearly a month now.特里由于功能性高烧已卧床近一个月了。
  • A malignant tumor originating in glandular tissue.腺癌起源于腺性组织的恶性肿瘤。
27 relinquished 2d789d1995a6a7f21bb35f6fc8d61c5d     
交出,让给( relinquish的过去式和过去分词 ); 放弃
参考例句:
  • She has relinquished the post to her cousin, Sir Edward. 她把职位让给了表弟爱德华爵士。
  • The small dog relinquished his bone to the big dog. 小狗把它的骨头让给那只大狗。
28 cerebral oUdyb     
adj.脑的,大脑的;有智力的,理智型的
参考例句:
  • Your left cerebral hemisphere controls the right-hand side of your body.你的左半脑控制身体的右半身。
  • He is a precise,methodical,cerebral man who carefully chooses his words.他是一个一丝不苟、有条理和理智的人,措辞谨慎。
29 volition cLkzS     
n.意志;决意
参考例句:
  • We like to think that everything we do and everything we think is a product of our volition.我们常常认为我们所做和所想的一切都出自自己的意愿。
  • Makin said Mr Coombes had gone to the police of his own volition.梅金说库姆斯先生是主动去投案的。
30 unravel Ajzwo     
v.弄清楚(秘密);拆开,解开,松开
参考例句:
  • He was good with his hands and could unravel a knot or untangle yarn that others wouldn't even attempt.他的手很灵巧,其他人甚至都不敢尝试的一些难解的绳结或缠在一起的纱线,他都能解开。
  • This is the attitude that led him to unravel a mystery that long puzzled Chinese historians.正是这种态度使他解决了长期以来使中国历史学家们大惑不解的谜。
31 physiological aAvyK     
adj.生理学的,生理学上的
参考例句:
  • He bought a physiological book.他买了一本生理学方面的书。
  • Every individual has a physiological requirement for each nutrient.每个人对每种营养成分都有一种生理上的需要。
32 reptile xBiz7     
n.爬行动物;两栖动物
参考例句:
  • The frog is not a true reptile.青蛙并非真正的爬行动物。
  • So you should not be surprised to see someone keep a reptile as a pet.所以,你不必惊奇有人养了一只爬行动物作为宠物。
33 essentially nntxw     
adv.本质上,实质上,基本上
参考例句:
  • Really great men are essentially modest.真正的伟人大都很谦虚。
  • She is an essentially selfish person.她本质上是个自私自利的人。
34 disturbance BsNxk     
n.动乱,骚动;打扰,干扰;(身心)失调
参考例句:
  • He is suffering an emotional disturbance.他的情绪受到了困扰。
  • You can work in here without any disturbance.在这儿你可不受任何干扰地工作。
35 psychical 8d18cc3bc74677380d4909fef11c68da     
adj.有关特异功能现象的;有关特异功能官能的;灵魂的;心灵的
参考例句:
  • Conclusion: The Liuhe-lottery does harm to people, s psychical health and should be for bidden. 结论:“六合彩”赌博有害人们心理卫生,应予以严禁。 来自互联网
36 instinctive c6jxT     
adj.(出于)本能的;直觉的;(出于)天性的
参考例句:
  • He tried to conceal his instinctive revulsion at the idea.他试图饰盖自己对这一想法本能的厌恶。
  • Animals have an instinctive fear of fire.动物本能地怕火。
37 preening 2d7802bbf088e82544268e2af08d571a     
v.(鸟)用嘴整理(羽毛)( preen的现在分词 )
参考例句:
  • Will you stop preening yourself in front of the mirror? 你别对着镜子打扮个没完行不行?
  • She was fading, while he was still preening himself in his elegance and youth. 她已显老,而他却仍然打扮成翩翩佳公子。 来自英汉文学 - 嘉莉妹妹
38 ledge o1Mxk     
n.壁架,架状突出物;岩架,岩礁
参考例句:
  • They paid out the line to lower him to the ledge.他们放出绳子使他降到那块岩石的突出部分。
  • Suddenly he struck his toe on a rocky ledge and fell.突然他的脚趾绊在一块突出的岩石上,摔倒了。
39 hesitation tdsz5     
n.犹豫,踌躇
参考例句:
  • After a long hesitation, he told the truth at last.踌躇了半天,他终于直说了。
  • There was a certain hesitation in her manner.她的态度有些犹豫不决。
40 beak 8y1zGA     
n.鸟嘴,茶壶嘴,钩形鼻
参考例句:
  • The bird had a worm in its beak.鸟儿嘴里叼着一条虫。
  • This bird employs its beak as a weapon.这种鸟用嘴作武器。
41 analogous aLdyQ     
adj.相似的;类似的
参考例句:
  • The two situations are roughly analogous.两种情況大致相似。
  • The company is in a position closely analogous to that of its main rival.该公司与主要竞争对手的处境极为相似。
42 noted 5n4zXc     
adj.著名的,知名的
参考例句:
  • The local hotel is noted for its good table.当地的那家酒店以餐食精美而著称。
  • Jim is noted for arriving late for work.吉姆上班迟到出了名。
43 neural DnXzFt     
adj.神经的,神经系统的
参考例句:
  • The neural network can preferably solve the non- linear problem.利用神经网络建模可以较好地解决非线性问题。
  • The information transmission in neural system depends on neurotransmitters.信息传递的神经途径有赖于神经递质。
44 apparatus ivTzx     
n.装置,器械;器具,设备
参考例句:
  • The school's audio apparatus includes films and records.学校的视听设备包括放映机和录音机。
  • They had a very refined apparatus.他们有一套非常精良的设备。
45 formerly ni3x9     
adv.从前,以前
参考例句:
  • We now enjoy these comforts of which formerly we had only heard.我们现在享受到了过去只是听说过的那些舒适条件。
  • This boat was formerly used on the rivers of China.这船从前航行在中国内河里。
46 secondly cjazXx     
adv.第二,其次
参考例句:
  • Secondly,use your own head and present your point of view.第二,动脑筋提出自己的见解。
  • Secondly it is necessary to define the applied load.其次,需要确定所作用的载荷。
47 stupor Kqqyx     
v.昏迷;不省人事
参考例句:
  • As the whisky took effect, he gradually fell into a drunken stupor.随着威士忌酒力发作,他逐渐醉得不省人事。
  • The noise of someone banging at the door roused her from her stupor.梆梆的敲门声把她从昏迷中唤醒了。
48 conspicuous spszE     
adj.明眼的,惹人注目的;炫耀的,摆阔气的
参考例句:
  • It is conspicuous that smoking is harmful to health.很明显,抽烟对健康有害。
  • Its colouring makes it highly conspicuous.它的色彩使它非常惹人注目。
49 phenomena 8N9xp     
n.现象
参考例句:
  • Ade couldn't relate the phenomena with any theory he knew.艾德无法用他所知道的任何理论来解释这种现象。
  • The object of these experiments was to find the connection,if any,between the two phenomena.这些实验的目的就是探索这两种现象之间的联系,如果存在着任何联系的话。
50 faculties 066198190456ba4e2b0a2bda2034dfc5     
n.能力( faculty的名词复数 );全体教职员;技巧;院
参考例句:
  • Although he's ninety, his mental faculties remain unimpaired. 他虽年届九旬,但头脑仍然清晰。
  • All your faculties have come into play in your work. 在你的工作中,你的全部才能已起到了作用。 来自《简明英汉词典》
51 mere rC1xE     
adj.纯粹的;仅仅,只不过
参考例句:
  • That is a mere repetition of what you said before.那不过是重复了你以前讲的话。
  • It's a mere waste of time waiting any longer.再等下去纯粹是浪费时间。
52 lateral 83ey7     
adj.侧面的,旁边的
参考例句:
  • An airfoil that controls lateral motion.能够控制横向飞行的机翼。
  • Mr.Dawson walked into the court from a lateral door.道森先生从一个侧面的门走进法庭。
53 lobes fe8c3178c8180f03dd0fc8ae16f13e3c     
n.耳垂( lobe的名词复数 );(器官的)叶;肺叶;脑叶
参考例句:
  • The rotor has recesses in its three faces between the lobes. 转子在其凸角之间的三个面上有凹槽。 来自辞典例句
  • The chalazal parts of the endosperm containing free nuclei forms several lobes. 包含游离核的合点端胚乳部分形成几个裂片。 来自辞典例句
54 affected TzUzg0     
adj.不自然的,假装的
参考例句:
  • She showed an affected interest in our subject.她假装对我们的课题感到兴趣。
  • His manners are affected.他的态度不自然。
55 vivacity ZhBw3     
n.快活,活泼,精神充沛
参考例句:
  • Her charm resides in her vivacity.她的魅力存在于她的活泼。
  • He was charmed by her vivacity and high spirits.她的活泼与兴高采烈的情绪把他迷住了。
56 physiologists c2a885ea249ea80fd0b5bfd528aedac0     
n.生理学者( physiologist的名词复数 );生理学( physiology的名词复数 );生理机能
参考例句:
  • Quite unexpectedly, vertebrate physiologists and microbial biochemists had found a common ground. 出乎意外,脊椎动物生理学家和微生物生化学家找到了共同阵地。 来自辞典例句
  • Physiologists are interested in the workings of the human body. 生理学家对人体的功能感兴趣。 来自辞典例句
57 respiration us7yt     
n.呼吸作用;一次呼吸;植物光合作用
参考例句:
  • They tried artificial respiration but it was of no avail.他们试做人工呼吸,可是无效。
  • They made frequent checks on his respiration,pulse and blood.他们经常检查他的呼吸、脉搏和血液。
58 irreconcilable 34RxO     
adj.(指人)难和解的,势不两立的
参考例句:
  • These practices are irreconcilable with the law of the Church.这种做法与教规是相悖的。
  • These old concepts are irreconcilable with modern life.这些陈旧的观念与现代生活格格不入。
59 mechanism zCWxr     
n.机械装置;机构,结构
参考例句:
  • The bones and muscles are parts of the mechanism of the body.骨骼和肌肉是人体的组成部件。
  • The mechanism of the machine is very complicated.这台机器的结构是非常复杂的。
60 supreme PHqzc     
adj.极度的,最重要的;至高的,最高的
参考例句:
  • It was the supreme moment in his life.那是他一生中最重要的时刻。
  • He handed up the indictment to the supreme court.他把起诉书送交最高法院。
61 allotted 5653ecda52c7b978bd6890054bd1f75f     
分配,拨给,摊派( allot的过去式和过去分词 )
参考例句:
  • I completed the test within the time allotted . 我在限定的时间内完成了试验。
  • Each passenger slept on the berth allotted to him. 每个旅客都睡在分配给他的铺位上。
62 mechanisms d0db71d70348ef1c49f05f59097917b8     
n.机械( mechanism的名词复数 );机械装置;[生物学] 机制;机械作用
参考例句:
  • The research will provide direct insight into molecular mechanisms. 这项研究将使人能够直接地了解分子的机理。 来自《简明英汉词典》
  • He explained how the two mechanisms worked. 他解释这两台机械装置是如何工作的。 来自《简明英汉词典》
63 interpretations a61815f6fe8955c9d235d4082e30896b     
n.解释( interpretation的名词复数 );表演;演绎;理解
参考例句:
  • This passage is open to a variety of interpretations. 这篇文章可以有各种不同的解释。 来自《简明英汉词典》
  • The involved and abstruse passage makes several interpretations possible. 这段艰涩的文字可以作出好几种解释。 来自《现代汉英综合大词典》
64 disappearance ouEx5     
n.消失,消散,失踪
参考例句:
  • He was hard put to it to explain her disappearance.他难以说明她为什么不见了。
  • Her disappearance gave rise to the wildest rumours.她失踪一事引起了各种流言蜚语。
65 persistence hSLzh     
n.坚持,持续,存留
参考例句:
  • The persistence of a cough in his daughter puzzled him.他女儿持续的咳嗽把他难住了。
  • He achieved success through dogged persistence.他靠着坚持不懈取得了成功。
66 chameleon YUWy2     
n.变色龙,蜥蜴;善变之人
参考例句:
  • The chameleon changes colour to match its surroundings.变色龙变换颜色以适应环境。
  • The chameleon can take on the colour of its background.变色龙可呈现出与其背景相同的颜色。
67 extirpation 24e80f0b67cdcaab1a1ccb18d37d9d8e     
n.消灭,根除,毁灭;摘除
参考例句:
  • Gamma Knife surgery has recently been tried as an alternative to surgical extirpation. 伽玛刀治疗最近被尝试作为手术根治之外的另一种选择。 来自辞典例句
  • Gamma Knife surgery (GKS) has recently been tried as an alternative to surgical extirpation. 伽玛刀治疗(GKS)最近被尝试作为手术根治之外的另一种选择。 来自互联网
68 biassed 6e85c46f87d4ad098e6df7e2de970b02     
(统计试验中)结果偏倚的,有偏的
参考例句:
69 persistently MlzztP     
ad.坚持地;固执地
参考例句:
  • He persistently asserted his right to a share in the heritage. 他始终声称他有分享那笔遗产的权利。
  • She persistently asserted her opinions. 她果断地说出了自己的意见。
70 jaws cq9zZq     
n.口部;嘴
参考例句:
  • The antelope could not escape the crocodile's gaping jaws. 那只羚羊无法从鱷鱼张开的大口中逃脱。
  • The scored jaws of a vise help it bite the work. 台钳上有刻痕的虎钳牙帮助它紧咬住工件。
71 vapors 94a2c1cb72b6aa4cb43b8fb8f61653d4     
n.水汽,水蒸气,无实质之物( vapor的名词复数 );自夸者;幻想 [药]吸入剂 [古]忧郁(症)v.自夸,(使)蒸发( vapor的第三人称单数 )
参考例句:
  • His emotions became vague and shifted about like vapors. 他的心情则如同一团雾气,变幻无常,捉摸不定。 来自辞典例句
  • They have hysterics, they weep, they have the vapors. 他们歇斯底里,他们哭泣,他们精神忧郁。 来自辞典例句
72 eyelid zlcxj     
n.眼睑,眼皮
参考例句:
  • She lifted one eyelid to see what he was doing.她抬起一只眼皮看看他在做什么。
  • My eyelid has been tumid since yesterday.从昨天起,我的眼皮就肿了。
73 hops a6b9236bf6c7a3dfafdbc0709208acc0     
跳上[下]( hop的第三人称单数 ); 单足蹦跳; 齐足(或双足)跳行; 摘葎草花
参考例句:
  • The sparrow crossed the lawn in a series of hops. 那麻雀一蹦一跳地穿过草坪。
  • It is brewed from malt and hops. 它用麦精和蛇麻草酿成。
74 remains 1kMzTy     
n.剩余物,残留物;遗体,遗迹
参考例句:
  • He ate the remains of food hungrily.他狼吞虎咽地吃剩余的食物。
  • The remains of the meal were fed to the dog.残羹剩饭喂狗了。
75 regain YkYzPd     
vt.重新获得,收复,恢复
参考例句:
  • He is making a bid to regain his World No.1 ranking.他正为重登世界排名第一位而努力。
  • The government is desperate to regain credibility with the public.政府急于重新获取公众的信任。
76 equilibrium jiazs     
n.平衡,均衡,相称,均势,平静
参考例句:
  • Change in the world around us disturbs our inner equilibrium.我们周围世界的变化扰乱了我们内心的平静。
  • This is best expressed in the form of an equilibrium constant.这最好用平衡常数的形式来表示。
77 variance MiXwb     
n.矛盾,不同
参考例句:
  • The question of woman suffrage sets them at variance. 妇女参政的问题使他们发生争执。
  • It is unnatural for brothers to be at variance. 兄弟之间不睦是不近人情的。
78 anatomy Cwgzh     
n.解剖学,解剖;功能,结构,组织
参考例句:
  • He found out a great deal about the anatomy of animals.在动物解剖学方面,他有过许多发现。
  • The hurricane's anatomy was powerful and complex.对飓风的剖析是一项庞大而复杂的工作。
79 acting czRzoc     
n.演戏,行为,假装;adj.代理的,临时的,演出用的
参考例句:
  • Ignore her,she's just acting.别理她,她只是假装的。
  • During the seventies,her acting career was in eclipse.在七十年代,她的表演生涯黯然失色。
80 volitional zh1wE     
adj.意志的,凭意志的,有意志的
参考例句:
  • The image consists in our rational,volitional,affective faculties,and in our bodies.神的形象存在于我们的理性、意志,和情感的能力中,也存在于我们的身体内。
  • The endurance of setbacks is a standard mat can evaluate the volitional character of a person.挫折承受能力是衡量人的意志品质的一个重要指标。
81 stimulation BuIwL     
n.刺激,激励,鼓舞
参考例句:
  • The playgroup provides plenty of stimulation for the children.幼儿游戏组给孩子很多启发。
  • You don't get any intellectual stimulation in this job.你不能从这份工作中获得任何智力启发。
82 dexterity hlXzs     
n.(手的)灵巧,灵活
参考例句:
  • You need manual dexterity to be good at video games.玩好电子游戏手要灵巧。
  • I'm your inferior in manual dexterity.论手巧,我不如你。
83 underlying 5fyz8c     
adj.在下面的,含蓄的,潜在的
参考例句:
  • The underlying theme of the novel is very serious.小说隐含的主题是十分严肃的。
  • This word has its underlying meaning.这个单词有它潜在的含义。
84 purely 8Sqxf     
adv.纯粹地,完全地
参考例句:
  • I helped him purely and simply out of friendship.我帮他纯粹是出于友情。
  • This disproves the theory that children are purely imitative.这证明认为儿童只会单纯地模仿的理论是站不住脚的。
85 distinguished wu9z3v     
adj.卓越的,杰出的,著名的
参考例句:
  • Elephants are distinguished from other animals by their long noses.大象以其长长的鼻子显示出与其他动物的不同。
  • A banquet was given in honor of the distinguished guests.宴会是为了向贵宾们致敬而举行的。
86 postponing 3ca610c0db966cd6f77cd5d15dc2b28c     
v.延期,推迟( postpone的现在分词 )
参考例句:
  • He tried to gain time by postponing his decision. 他想以迟迟不作决定的手段来争取时间。 来自辞典例句
  • I don't hold with the idea of postponing further discussion of the matter. 我不赞成推迟进一步讨论这件事的想法。 来自辞典例句
87 doctrine Pkszt     
n.教义;主义;学说
参考例句:
  • He was impelled to proclaim his doctrine.他不得不宣扬他的教义。
  • The council met to consider changes to doctrine.宗教议会开会考虑更改教义。


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