As to the oviparous quadrupeds, and such birds as resemble them in closing the eye with the lower lid, it is the hardness of the skin of their heads which makes them do so. For such birds as have heavy bodies are not made for flight; and so the materials which would otherwise have gone to increase the growth of the feathers are diverted thence, and used to augment4 the thickness of the skin. Birds therefore of this kind close the eye with the lower lid; whereas pigeons and the like use both upper and lower lids for the purpose. As birds are covered with feathers, so oviparous quadrupeds are covered with scaly5 plates; and these in all their forms are harder than hairs, so that the skin also to which they belong is harder than the skin of hairy animals. In these animals, then, the skin on the head is hard, and so does not allow of the formation of an upper eyelid6, whereas lower down the integument7 is of a flesh-like character, so that the lower lid can be thin and extensible.
The act of blinking is performed by the heavy-bodied birds by means of the membrane already mentioned, and not by this lower lid. For in blinking rapid motion is required, and such is the motion of this membrane, whereas that of the lower lid is slow. It is from the canthus that is nearest to the nostrils8 that the membrane comes. For it is better to have one starting-point for nictitation than two; and in these birds this starting-point is the junction9 of eye and nostrils, an anterior10 starting-point being preferable to a lateral11 one. Oviparous quadrupeds do not blink in like manner as the birds; for, living as they do on the ground, they are free from the necessity of having eyes of fluid consistency and of keen sight, whereas these are essential requisites12 for birds, inasmuch as they have to use their eyes at long distances. This too explains why birds with talons13, that have to search for prey14 by eye from aloft, and therefore soar to greater heights than other birds, are sharpsighted; while common fowls15 and the like, that live on the ground and are not made for flight, have no such keenness of vision. For there is nothing in their mode of life which imperatively16 requires it.
Fishes and Insects and the hard-skinned Crustacea present certain differences in their eyes, but so far resemble each other as that none of them have eyelids17. As for the hard-skinned Crustacea it is utterly18 out of the question that they should have any; for an eyelid, to be of use, requires the action of the skin to be rapid. These animals then have no eyelids and, in default of this protection, their eyes are hard, just as though the lid were attached to the surface of the eye, and the animal saw through it. Inasmuch, however, as such hardness must necessarily blunt the sharpness of vision, nature has endowed the eyes of Insects, and still more those of Crustacea, with mobility19 (just as she has given some quadrupeds movable ears), in order that they may be able to turn to the light and catch its rays, and so see more plainly. Fishes, however, have eyes of a fluid consistency. For animals that move much about have to use their vision at considerable distances. If now they live on land, the air in which they move is transparent20 enough. But the water in which fishes live is a hindrance21 to sharp sight, though it has this advantage over the air, that it does not contain so many objects to knock against the eyes. The risk of collision being thus small, nature, who makes nothing in vain, has given no eyelids to fishes, while to counterbalance the opacity22 of the water she has made their eyes of fluid consistency.
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1 membrane | |
n.薄膜,膜皮,羊皮纸 | |
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2 consistency | |
n.一贯性,前后一致,稳定性;(液体的)浓度 | |
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3 delicacy | |
n.精致,细微,微妙,精良;美味,佳肴 | |
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4 augment | |
vt.(使)增大,增加,增长,扩张 | |
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5 scaly | |
adj.鱼鳞状的;干燥粗糙的 | |
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6 eyelid | |
n.眼睑,眼皮 | |
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7 integument | |
n.皮肤 | |
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8 nostrils | |
鼻孔( nostril的名词复数 ) | |
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9 junction | |
n.连接,接合;交叉点,接合处,枢纽站 | |
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10 anterior | |
adj.较早的;在前的 | |
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11 lateral | |
adj.侧面的,旁边的 | |
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12 requisites | |
n.必要的事物( requisite的名词复数 ) | |
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13 talons | |
n.(尤指猛禽的)爪( talon的名词复数 );(如爪般的)手指;爪状物;锁簧尖状突出部 | |
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14 prey | |
n.被掠食者,牺牲者,掠食;v.捕食,掠夺,折磨 | |
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15 fowls | |
鸟( fowl的名词复数 ); 禽肉; 既不是这; 非驴非马 | |
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16 imperatively | |
adv.命令式地 | |
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17 eyelids | |
n.眼睑( eyelid的名词复数 );眼睛也不眨一下;不露声色;面不改色 | |
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18 utterly | |
adv.完全地,绝对地 | |
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19 mobility | |
n.可动性,变动性,情感不定 | |
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20 transparent | |
adj.明显的,无疑的;透明的 | |
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21 hindrance | |
n.妨碍,障碍 | |
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22 opacity | |
n.不透明;难懂 | |
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