Birds also present variations in the part which acts as a recipient23 of the food; and the reason for these variations is the same as in the animals just mentioned. For here again it is because the mouth fails to perform its office and fails even more completely-for birds have no teeth at all, nor any instrument whatsoever24 with which to comminute or grind down their food-it is, I say, because of this, that in some of them what is called the crop precedes the stomach and does the work of the mouth; while in others the oesophagus is either wide throughout or a part of it bulges25 just before it enters the stomach, so as to form a preparatory store-house for the unreduced food; or the stomach itself has a protuberance in some part, or is strong and fleshy, so as to be able to store up the food for a considerable period and to concoct6 it, in spite of its not having been ground into a pulp. For nature retrieves26 the inefficiency27 of the mouth by increasing the efficiency and heat of the stomach. Other birds there are, such, namely, as have long legs and live in marshes28, that have none of these provisions, but merely an elongated29 oesophagus. The explanation of this is to be found in the moist character of their food. For all these birds feed on substances easy of reduction, and their food being moist and not requiring much concoction, their digestive cavities are of a corresponding character.
Fishes are provided with teeth, which in almost all of them are of the sharp interfitting kind. For there is but one small section in which it is otherwise. Of these the fish called Scarus (Parrot-fish) is an example. And this is probably the reason why this fish apparently30 ruminates, though no other fishes do so. For those horned animals that have no front teeth in the upper jaw also ruminate17.
In fishes the teeth are all sharp; so that these animals can divide their food, though imperfectly. For it is impossible for a fish to linger or spend time in the act of mastication31, and therefore they have no teeth that are flat or suitable for grinding; for such teeth would be to no purpose. The oesophagus again in some fishes is entirely32 wanting, and in the rest is but short. In order, however, to facilitate the concoction of the food, some of them, as the Cestreus (mullet), have a fleshy stomach resembling that of a bird; while most of them have numerous processes close against the stomach, to serve as a sort of antechamber in which the food may be stored up and undergo putrefaction33 and concoction. There is contrast between fishes and birds in the position of these processes. For in fishes they are placed close to the stomach; while in birds, if present at all, they are lower down, near the end of the gut. Some of the Vivipara also have processes connected with the lower part of the gut which serve the same purpose as that stated above.
The whole tribe of fishes is of gluttonous34 appetite, owing to the arrangements for the reduction of their food being very imperfect, and much of it consequently passing through them without undergoing concoction; and, of all, those are the most gluttonous that have a straight intestine35. For as the passage of food in such cases is rapid, and the enjoyment36 derived37 from it in consequence but brief, it follows of necessity that the return of appetite is also speedy.
It has already been mentioned that in animals with front teeth in both jaws the stomach is of small size. It may be classed pretty nearly always under one or other of two headings, namely as resembling the stomach of the dog, or as resembling the stomach of the pig. In the pig the stomach is larger than in the dog, and presents certain folds of moderate size, the purpose of which is to lengthen38 out the period of concoction; while the stomach of the dog is of small size, not much larger in calibre than the gut, and smooth on the internal surface.
Not much larger, I say, than the gut; for in all animals after the stomach comes the gut. This, like the stomach, presents numerous modifications39. For in some animals it is uniform, when uncoiled, and alike throughout, while in others it differs in different portions. Thus in some cases it is wider in the neighbourhood of the stomach, and narrower towards the other end; and this explains by the way why dogs have to strain so much in discharging their excrement40. But in most animals it is the upper portion that is the narrower and the lower that is of greater width.
Of greater length than in other animals, and much convoluted41, are the intestines42 of those that have horns. These intestines, moreover, as also the stomach, are of ampler volume, in accordance with the larger size of the body. For animals with horns are, as a rule, animals of no small bulk, because of the thorough elaboration which their food undergoes. The gut, except in those animals where it is straight, invariably widens out as we get farther from the stomach and come to what is called the colon43, and to a kind of caecal dilatation. After this it again becomes narrower and convoluted. Then succeeds a straight portion which runs right on to the vent44. This vent is known as the anus, and is in some animals surrounded by fat, in others not so. All these parts have been so contrived45 by nature as to harmonize with the various operations that relate to the food and its residue. For, as the residual food gets farther on and lower down, the space to contain it enlarges, allowing it to remain stationary46 and undergo conversion47. Thus is it in those animals which, owing either to their large size, or to the heat of the parts concerned, require more nutriment, and consume more fodder48 than the rest.
Neither is it without a purpose, that, just as a narrower gut succeeds to the upper stomach, so also does the residual food, when its goodness is thoroughly49 exhausted, pass from the colon and the ample space of the lower stomach into a narrower channel and into the spiral coil. For so nature can regulate her expenditure50 and prevent the excremental51 residue from being discharged all at once.
In all such animals, however, as have to be comparatively moderate in their alimentation, the lower stomach presents no wide and roomy spaces, though their gut is not straight, but has a number of convolutions. For amplitude52 of space causes desire for ample food, and straightness of the intestine causes quick return of appetite. And thus it is that all animals whose food receptacles are either simple or spacious53 are of gluttonous habits, the latter eating enormously at a meal, the former making meals at short intervals54.
Again, since the food in the upper stomach, having just been swallowed, must of necessity be quite fresh, while that which has reached the lower stomach must have had its juices exhausted and resemble dung, it follows of necessity that there must also be some intermediate part, in which the change may be effected, and where the food will be neither perfectly19 fresh nor yet dung. And thus it is that, in all such animals as we are now considering, there is found what is called the jejunum; which is a part of the small gut, of the gut, that is, which comes next to the stomach. For this jejunum lies between the upper cavity which contains the yet unconcocted food and the lower cavity which holds the residual matter, which by the time it has got here has become worthless. There is a jejunum in all these animals, but it is only plainly discernible in those of large size, and this only when they have abstained55 from food for a certain time. For then alone can one hit on the exact period when the food lies half-way between the upper and lower cavities; a period which is very short, for the time occupied in the transition of food is but brief. In females this jejunum may occupy any part whatsoever of the upper intestine, but in males it comes just before the caecum and the lower stomach.
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1 immediate | |
adj.立即的;直接的,最接近的;紧靠的 | |
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2 contiguity | |
n.邻近,接壤 | |
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3 gut | |
n.[pl.]胆量;内脏;adj.本能的;vt.取出内脏 | |
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4 exhausted | |
adj.极其疲惫的,精疲力尽的 | |
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5 residual | |
adj.复播复映追加时间;存留下来的,剩余的 | |
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6 concoct | |
v.调合,制造 | |
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7 residue | |
n.残余,剩余,残渣 | |
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8 forth | |
adv.向前;向外,往外 | |
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9 jaws | |
n.口部;嘴 | |
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10 jaw | |
n.颚,颌,说教,流言蜚语;v.喋喋不休,教训 | |
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11 hoof | |
n.(马,牛等的)蹄 | |
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12 mule | |
n.骡子,杂种,执拗的人 | |
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13 ass | |
n.驴;傻瓜,蠢笨的人 | |
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14 thorny | |
adj.多刺的,棘手的 | |
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15 ligneous | |
adj.木质的,木头的 | |
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16 concoction | |
n.调配(物);谎言 | |
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17 ruminate | |
v.反刍;沉思 | |
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18 ruminates | |
v.沉思( ruminate的第三人称单数 );反复考虑;反刍;倒嚼 | |
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19 perfectly | |
adv.完美地,无可非议地,彻底地 | |
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20 pulp | |
n.果肉,纸浆;v.化成纸浆,除去...果肉,制成纸浆 | |
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21 treatises | |
n.专题著作,专题论文,专著( treatise的名词复数 ) | |
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22 anatomy | |
n.解剖学,解剖;功能,结构,组织 | |
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23 recipient | |
a.接受的,感受性强的 n.接受者,感受者,容器 | |
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24 whatsoever | |
adv.(用于否定句中以加强语气)任何;pron.无论什么 | |
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25 bulges | |
膨胀( bulge的名词复数 ); 鼓起; (身体的)肥胖部位; 暂时的激增 | |
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26 retrieves | |
v.取回( retrieve的第三人称单数 );恢复;寻回;检索(储存的信息) | |
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27 inefficiency | |
n.无效率,无能;无效率事例 | |
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28 marshes | |
n.沼泽,湿地( marsh的名词复数 ) | |
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29 elongated | |
v.延长,加长( elongate的过去式和过去分词 ) | |
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30 apparently | |
adv.显然地;表面上,似乎 | |
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31 mastication | |
n.咀嚼 | |
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32 entirely | |
ad.全部地,完整地;完全地,彻底地 | |
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33 putrefaction | |
n.腐坏,腐败 | |
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34 gluttonous | |
adj.贪吃的,贪婪的 | |
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35 intestine | |
adj.内部的;国内的;n.肠 | |
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36 enjoyment | |
n.乐趣;享有;享用 | |
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37 derived | |
vi.起源;由来;衍生;导出v.得到( derive的过去式和过去分词 );(从…中)得到获得;源于;(从…中)提取 | |
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38 lengthen | |
vt.使伸长,延长 | |
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39 modifications | |
n.缓和( modification的名词复数 );限制;更改;改变 | |
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40 excrement | |
n.排泄物,粪便 | |
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41 convoluted | |
adj.旋绕的;复杂的 | |
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42 intestines | |
n.肠( intestine的名词复数 ) | |
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43 colon | |
n.冒号,结肠,直肠 | |
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44 vent | |
n.通风口,排放口;开衩;vt.表达,发泄 | |
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45 contrived | |
adj.不自然的,做作的;虚构的 | |
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46 stationary | |
adj.固定的,静止不动的 | |
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47 conversion | |
n.转化,转换,转变 | |
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48 fodder | |
n.草料;炮灰 | |
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49 thoroughly | |
adv.完全地,彻底地,十足地 | |
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50 expenditure | |
n.(时间、劳力、金钱等)支出;使用,消耗 | |
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51 excremental | |
adj.排泄物的,粪便的 | |
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52 amplitude | |
n.广大;充足;振幅 | |
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53 spacious | |
adj.广阔的,宽敞的 | |
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54 intervals | |
n.[军事]间隔( interval的名词复数 );间隔时间;[数学]区间;(戏剧、电影或音乐会的)幕间休息 | |
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55 abstained | |
v.戒(尤指酒),戒除( abstain的过去式和过去分词 );弃权(不投票) | |
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