On the absence of intermediate varieties at the present day. On the nature of extinct intermediate varieties; on their number. On the vast lapse1 of time, as inferred from the rate of deposition2 and of denudation3. On the poorness of our palaeontological collections. On the intermittence4 of geological formations. On the absence of intermediate varieties in any one formation. On the sudden appearance of groups of species. On their sudden appearance in the lowest known fossiliferous strata5.
In the sixth chapter I enumerated6 the chief objections which might be justly urged against the views maintained in this volume. Most of them have now been discussed. One, namely the distinctness of specific forms, and their not being blended together by innumerable transitional links, is a very obvious difficulty. I assigned reasons why such links do not commonly occur at the present day, under the circumstances apparently7 most favourable8 for their presence, namely on an extensive and continuous area with graduated physical conditions. I endeavoured to show, that the life of each species depends in a more important manner on the presence of other already defined organic forms, than on climate; and, therefore, that the really governing conditions of life do not graduate away quite insensibly like heat or moisture. I endeavoured, also, to show that intermediate varieties, from existing in lesser9 numbers than the forms which they connect, will generally be beaten out and exterminated10 during the course of further modification12 and improvement. The main cause, however, of innumerable intermediate links not now occurring everywhere throughout nature depends on the very process of natural selection, through which new varieties continually take the places of and exterminate11 their parent-forms. But just in proportion as this process of extermination13 has acted on an enormous scale, so must the number of intermediate varieties, which have formerly14 existed on the earth, be truly enormous. Why then is not every geological formation and every stratum15 full of such intermediate links? Geology assuredly does not reveal any such finely graduated organic chain; and this, perhaps, is the most obvious and gravest objection which can be urged against my theory. The explanation lies, as I believe, in the extreme imperfection of the geological record.
In the first place it should always be borne in mind what sort of intermediate forms must, on my theory, have formerly existed. I have found it difficult, when looking at any two species, to avoid picturing to myself, forms DIRECTLY intermediate between them. But this is a wholly false view; we should always look for forms intermediate between each species and a common but unknown progenitor16; and the progenitor will generally have differed in some respects from all its modified descendants. To give a simple illustration: the fantail and pouter pigeons have both descended17 from the rock-pigeon; if we possessed18 all the intermediate varieties which have ever existed, we should have an extremely close series between both and the rock-pigeon; but we should have no varieties directly intermediate between the fantail and pouter; none, for instance, combining a tail somewhat expanded with a crop somewhat enlarged, the characteristic features of these two breeds. These two breeds, moreover, have become so much modified, that if we had no historical or indirect evidence regarding their origin, it would not have been possible to have determined19 from a mere20 comparison of their structure with that of the rock-pigeon, whether they had descended from this species or from some other allied21 species, such as C. oenas.
So with natural species, if we look to forms very distinct, for instance to the horse and tapir, we have no reason to suppose that links ever existed directly intermediate between them, but between each and an unknown common parent. The common parent will have had in its whole organisation22 much general resemblance to the tapir and to the horse; but in some points of structure may have differed considerably23 from both, even perhaps more than they differ from each other. Hence in all such cases, we should be unable to recognise the parent-form of any two or more species, even if we closely compared the structure of the parent with that of its modified descendants, unless at the same time we had a nearly perfect chain of the intermediate links.
It is just possible by my theory, that one of two living forms might have descended from the other; for instance, a horse from a tapir; and in this case DIRECT intermediate links will have existed between them. But such a case would imply that one form had remained for a very long period unaltered, whilst its descendants had undergone a vast amount of change; and the principle of competition between organism and organism, between child and parent, will render this a very rare event; for in all cases the new and improved forms of life will tend to supplant24 the old and unimproved forms.
By the theory of natural selection all living species have been connected with the parent-species of each genus, by differences not greater than we see between the varieties of the same species at the present day; and these parent-species, now generally extinct, have in their turn been similarly connected with more ancient species; and so on backwards25, always converging26 to the common ancestor of each great class. So that the number of intermediate and transitional links, between all living and extinct species, must have been inconceivably great. But assuredly, if this theory be true, such have lived upon this earth.
ON THE LAPSE OF TIME.
Independently of our not finding fossil remains27 of such infinitely28 numerous connecting links, it may be objected, that time will not have sufficed for so great an amount of organic change, all changes having been effected very slowly through natural selection. It is hardly possible for me even to recall to the reader, who may not be a practical geologist29, the facts leading the mind feebly to comprehend the lapse of time. He who can read Sir Charles Lyell's grand work on the Principles of Geology, which the future historian will recognise as having produced a revolution in natural science, yet does not admit how incomprehensibly vast have been the past periods of time, may at once close this volume. Not that it suffices to study the Principles of Geology, or to read special treatises30 by different observers on separate formations, and to mark how each author attempts to give an inadequate31 idea of the duration of each formation or even each stratum. A man must for years examine for himself great piles of superimposed strata, and watch the sea at work grinding down old rocks and making fresh sediment32, before he can hope to comprehend anything of the lapse of time, the monuments of which we see around us.
It is good to wander along lines of sea-coast, when formed of moderately hard rocks, and mark the process of degradation33. The tides in most cases reach the cliffs only for a short time twice a day, and the waves eat into them only when they are charged with sand or pebbles34; for there is reason to believe that pure water can effect little or nothing in wearing away rock. At last the base of the cliff is undermined, huge fragments fall down, and these remaining fixed35, have to be worn away, atom by atom, until reduced in size they can be rolled about by the waves, and then are more quickly ground into pebbles, sand, or mud. But how often do we see along the bases of retreating cliffs rounded boulders36, all thickly clothed by marine37 productions, showing how little they are abraded38 and how seldom they are rolled about! Moreover, if we follow for a few miles any line of rocky cliff, which is undergoing degradation, we find that it is only here and there, along a short length or round a promontory39, that the cliffs are at the present time suffering. The appearance of the surface and the vegetation show that elsewhere years have elapsed since the waters washed their base.
He who most closely studies the action of the sea on our shores, will, I believe, be most deeply impressed with the slowness with which rocky coasts are worn away. The observations on this head by Hugh Miller40, and by that excellent observer Mr. Smith of Jordan Hill, are most impressive. With the mind thus impressed, let any one examine beds of conglomerate41 many thousand feet in thickness, which, though probably formed at a quicker rate than many other deposits, yet, from being formed of worn and rounded pebbles, each of which bears the stamp of time, are good to show how slowly the mass has been accumulated. Let him remember Lyell's profound remark, that the thickness and extent of sedimentary formations are the result and measure of the degradation which the earth's crust has elsewhere suffered. And what an amount of degradation is implied by the sedimentary deposits of many countries! Professor Ramsay has given me the maximum thickness, in most cases from actual measurement, in a few cases from estimate, of each formation in different parts of Great Britain; and this is the result:--
Feet Palaeozoic strata (not including igneous42 beds)..57,154. Secondary strata................................13,190. Tertiary strata..................................2,240.
--making altogether 72,584 feet; that is, very nearly thirteen and three-quarters British miles. Some of these formations, which are represented in England by thin beds, are thousands of feet in thickness on the Continent. Moreover, between each successive formation, we have, in the opinion of most geologists43, enormously long blank periods. So that the lofty pile of sedimentary rocks in Britain, gives but an inadequate idea of the time which has elapsed during their accumulation; yet what time this must have consumed! Good observers have estimated that sediment is deposited by the great Mississippi river at the rate of only 600 feet in a hundred thousand years. This estimate may be quite erroneous; yet, considering over what wide spaces very fine sediment is transported by the currents of the sea, the process of accumulation in any one area must be extremely slow.
But the amount of denudation which the strata have in many places suffered, independently of the rate of accumulation of the degraded matter, probably offers the best evidence of the lapse of time. I remember having been much struck with the evidence of denudation, when viewing volcanic44 islands, which have been worn by the waves and pared all round into perpendicular45 cliffs of one or two thousand feet in height; for the gentle slope of the lava-streams, due to their formerly liquid state, showed at a glance how far the hard, rocky beds had once extended into the open ocean. The same story is still more plainly told by faults,--those great cracks along which the strata have been upheaved on one side, or thrown down on the other, to the height or depth of thousands of feet; for since the crust cracked, the surface of the land has been so completely planed down by the action of the sea, that no trace of these vast dislocations is externally visible.
The Craven fault, for instance, extends for upwards46 of 30 miles, and along this line the vertical47 displacement48 of the strata has varied49 from 600 to 3000 feet. Professor Ramsay has published an account of a downthrow in Anglesea of 2300 feet; and he informs me that he fully50 believes there is one in Merionethshire of 12,000 feet; yet in these cases there is nothing on the surface to show such prodigious51 movements; the pile of rocks on the one or other side having been smoothly52 swept away. The consideration of these facts impresses my mind almost in the same manner as does the vain endeavour to grapple with the idea of eternity53.
I am tempted54 to give one other case, the well-known one of the denudation of the Weald. Though it must be admitted that the denudation of the Weald has been a mere trifle, in comparison with that which has removed masses of our palaeozoic strata, in parts ten thousand feet in thickness, as shown in Professor Ramsay's masterly memoir55 on this subject. Yet it is an admirable lesson to stand on the North Downs and to look at the distant South Downs; for, remembering that at no great distance to the west the northern and southern escarpments meet and close, one can safely picture to oneself the great dome56 of rocks which must have covered up the Weald within so limited a period as since the latter part of the Chalk formation. The distance from the northern to the southern Downs is about 22 miles, and the thickness of the several formations is on an average about 1100 feet, as I am informed by Professor Ramsay. But if, as some geologists suppose, a range of older rocks underlies57 the Weald, on the flanks of which the overlying sedimentary deposits might have accumulated in thinner masses than elsewhere, the above estimate would be erroneous; but this source of doubt probably would not greatly affect the estimate as applied58 to the western extremity59 of the district. If, then, we knew the rate at which the sea commonly wears away a line of cliff of any given height, we could measure the time requisite60 to have denuded61 the Weald. This, of course, cannot be done; but we may, in order to form some crude notion on the subject, assume that the sea would eat into cliffs 500 feet in height at the rate of one inch in a century. This will at first appear much too small an allowance; but it is the same as if we were to assume a cliff one yard in height to be eaten back along a whole line of coast at the rate of one yard in nearly every twenty-two years. I doubt whether any rock, even as soft as chalk, would yield at this rate excepting on the most exposed coasts; though no doubt the degradation of a lofty cliff would be more rapid from the breakage of the fallen fragments. On the other hand, I do not believe that any line of coast, ten or twenty miles in length, ever suffers degradation at the same time along its whole indented62 length; and we must remember that almost all strata contain harder layers or nodules, which from long resisting attrition form a breakwater at the base. Hence, under ordinary circumstances, I conclude that for a cliff 500 feet in height, a denudation of one inch per century for the whole length would be an ample allowance. At this rate, on the above data, the denudation of the Weald must have required 306,662,400 years; or say three hundred million years.
The action of fresh water on the gently inclined Wealden district, when upraised, could hardly have been great, but it would somewhat reduce the above estimate. On the other hand, during oscillations of level, which we know this area has undergone, the surface may have existed for millions of years as land, and thus have escaped the action of the sea: when deeply submerged for perhaps equally long periods, it would, likewise, have escaped the action of the coast-waves. So that in all probability a far longer period than 300 million years has elapsed since the latter part of the Secondary period.
I have made these few remarks because it is highly important for us to gain some notion, however imperfect, of the lapse of years. During each of these years, over the whole world, the land and the water has been peopled by hosts of living forms. What an infinite number of generations, which the mind cannot grasp, must have succeeded each other in the long roll of years! Now turn to our richest geological museums, and what a paltry64 display we behold65!
ON THE POORNESS OF OUR PALAEONTOLOGICAL COLLECTIONS.
That our palaeontological collections are very imperfect, is admitted by every one. The remark of that admirable palaeontologist, the late Edward Forbes, should not be forgotten, namely, that numbers of our fossil species are known and named from single and often broken specimens67, or from a few specimens collected on some one spot. Only a small portion of the surface of the earth has been geologically explored, and no part with sufficient care, as the important discoveries made every year in Europe prove. No organism wholly soft can be preserved. Shells and bones will decay and disappear when left on the bottom of the sea, where sediment is not accumulating. I believe we are continually taking a most erroneous view, when we tacitly admit to ourselves that sediment is being deposited over nearly the whole bed of the sea, at a rate sufficiently68 quick to embed69 and preserve fossil remains. Throughout an enormously large proportion of the ocean, the bright blue tint70 of the water bespeaks71 its purity. The many cases on record of a formation conformably covered, after an enormous interval72 of time, by another and later formation, without the underlying73 bed having suffered in the interval any wear and tear, seem explicable only on the view of the bottom of the sea not rarely lying for ages in an unaltered condition. The remains which do become embedded74, if in sand or gravel75, will when the beds are upraised generally be dissolved by the percolation76 of rain-water. I suspect that but few of the very many animals which live on the beach between high and low watermark are preserved. For instance, the several species of the Chthamalinae (a sub-family of sessile cirripedes) coat the rocks all over the world in infinite numbers: they are all strictly77 littoral78, with the exception of a single Mediterranean79 species, which inhabits deep water and has been found fossil in Sicily, whereas not one other species has hitherto been found in any tertiary formation: yet it is now known that the genus Chthamalus existed during the chalk period. The molluscan genus Chiton offers a partially80 analogous81 case.
With respect to the terrestrial productions which lived during the Secondary and Palaeozoic periods, it is superfluous82 to state that our evidence from fossil remains is fragmentary in an extreme degree. For instance, not a land shell is known belonging to either of these vast periods, with one exception discovered by Sir C. Lyell in the carboniferous strata of North America. In regard to mammiferous remains, a single glance at the historical table published in the Supplement to Lyell's Manual, will bring home the truth, how accidental and rare is their preservation83, far better than pages of detail. Nor is their rarity surprising, when we remember how large a proportion of the bones of tertiary mammals have been discovered either in caves or in lacustrine deposits; and that not a cave or true lacustrine bed is known belonging to the age of our secondary or palaeozoic formations.
But the imperfection in the geological record mainly results from another and more important cause than any of the foregoing; namely, from the several formations being separated from each other by wide intervals84 of time. When we see the formations tabulated85 in written works, or when we follow them in nature, it is difficult to avoid believing that they are closely consecutive86. But we know, for instance, from Sir R. Murchison's great work on Russia, what wide gaps there are in that country between the superimposed formations; so it is in North America, and in many other parts of the world. The most skilful87 geologist, if his attention had been exclusively confined to these large territories, would never have suspected that during the periods which were blank and barren in his own country, great piles of sediment, charged with new and peculiar88 forms of life, had elsewhere been accumulated. And if in each separate territory, hardly any idea can be formed of the length of time which has elapsed between the consecutive formations, we may infer that this could nowhere be ascertained89. The frequent and great changes in the mineralogical composition of consecutive formations, generally implying great changes in the geography of the surrounding lands, whence the sediment has been derived90, accords with the belief of vast intervals of time having elapsed between each formation.
But we can, I think, see why the geological formations of each region are almost invariably intermittent91; that is, have not followed each other in close sequence. Scarcely any fact struck me more when examining many hundred miles of the South American coasts, which have been upraised several hundred feet within the recent period, than the absence of any recent deposits sufficiently extensive to last for even a short geological period. Along the whole west coast, which is inhabited by a peculiar marine fauna92, tertiary beds are so scantily93 developed, that no record of several successive and peculiar marine faunas94 will probably be preserved to a distant age. A little reflection will explain why along the rising coast of the western side of South America, no extensive formations with recent or tertiary remains can anywhere be found, though the supply of sediment must for ages have been great, from the enormous degradation of the coast-rocks and from muddy streams entering the sea. The explanation, no doubt, is, that the littoral and sub-littoral deposits are continually worn away, as soon as they are brought up by the slow and gradual rising of the land within the grinding action of the coast-waves.
We may, I think, safely conclude that sediment must be accumulated in extremely thick, solid, or extensive masses, in order to withstand the incessant95 action of the waves, when first upraised and during subsequent oscillations of level. Such thick and extensive accumulations of sediment may be formed in two ways; either, in profound depths of the sea, in which case, judging from the researches of E. Forbes, we may conclude that the bottom will be inhabited by extremely few animals, and the mass when upraised will give a most imperfect record of the forms of life which then existed; or, sediment may be accumulated to any thickness and extent over a shallow bottom, if it continue slowly to subside96. In this latter case, as long as the rate of subsidence and supply of sediment nearly balance each other, the sea will remain shallow and favourable for life, and thus a fossiliferous formation thick enough, when upraised, to resist any amount of degradation, may be formed.
I am convinced that all our ancient formations, which are rich in fossils, have thus been formed during subsidence. Since publishing my views on this subject in 1845, I have watched the progress of Geology, and have been surprised to note how author after author, in treating of this or that great formation, has come to the conclusion that it was accumulated during subsidence. I may add, that the only ancient tertiary formation on the west coast of South America, which has been bulky enough to resist such degradation as it has as yet suffered, but which will hardly last to a distant geological age, was certainly deposited during a downward oscillation of level, and thus gained considerable thickness.
All geological facts tell us plainly that each area has undergone numerous slow oscillations of level, and apparently these oscillations have affected97 wide spaces. Consequently formations rich in fossils and sufficiently thick and extensive to resist subsequent degradation, may have been formed over wide spaces during periods of subsidence, but only where the supply of sediment was sufficient to keep the sea shallow and to embed and preserve the remains before they had time to decay. On the other hand, as long as the bed of the sea remained stationary98, THICK deposits could not have been accumulated in the shallow parts, which are the most favourable to life. Still less could this have happened during the alternate periods of elevation99; or, to speak more accurately100, the beds which were then accumulated will have been destroyed by being upraised and brought within the limits of the coast-action.
Thus the geological record will almost necessarily be rendered intermittent. I feel much confidence in the truth of these views, for they are in strict accordance with the general principles inculcated by Sir C. Lyell; and E. Forbes independently arrived at a similar conclusion.
One remark is here worth a passing notice. During periods of elevation the area of the land and of the adjoining shoal parts of the sea will be increased, and new stations will often be formed;--all circumstances most favourable, as previously101 explained, for the formation of new varieties and species; but during such periods there will generally be a blank in the geological record. On the other hand, during subsidence, the inhabited area and number of inhabitants will decrease (excepting the productions on the shores of a continent when first broken up into an archipelago), and consequently during subsidence, though there will be much extinction102, fewer new varieties or species will be formed; and it is during these very periods of subsidence, that our great deposits rich in fossils have been accumulated. Nature may almost be said to have guarded against the frequent discovery of her transitional or linking forms.
From the foregoing considerations it cannot be doubted that the geological record, viewed as a whole, is extremely imperfect; but if we confine our attention to any one formation, it becomes more difficult to understand, why we do not therein find closely graduated varieties between the allied species which lived at its commencement and at its close. Some cases are on record of the same species presenting distinct varieties in the upper and lower parts of the same formation, but, as they are rare, they may be here passed over. Although each formation has indisputably required a vast number of years for its deposition, I can see several reasons why each should not include a graduated series of links between the species which then lived; but I can by no means pretend to assign due proportional weight to the following considerations.
Although each formation may mark a very long lapse of years, each perhaps is short compared with the period requisite to change one species into another. I am aware that two palaeontologists, whose opinions are worthy103 of much deference104, namely Bronn and Woodward, have concluded that the average duration of each formation is twice or thrice as long as the average duration of specific forms. But insuperable difficulties, as it seems to me, prevent us coming to any just conclusion on this head. When we see a species first appearing in the middle of any formation, it would be rash in the extreme to infer that it had not elsewhere previously existed. So again when we find a species disappearing before the uppermost layers have been deposited, it would be equally rash to suppose that it then became wholly extinct. We forget how small the area of Europe is compared with the rest of the world; nor have the several stages of the same formation throughout Europe been correlated with perfect accuracy.
With marine animals of all kinds, we may safely infer a large amount of migration105 during climatal and other changes; and when we see a species first appearing in any formation, the probability is that it only then first immigrated106 into that area. It is well known, for instance, that several species appeared somewhat earlier in the palaeozoic beds of North America than in those of Europe; time having apparently been required for their migration from the American to the European seas. In examining the latest deposits of various quarters of the world, it has everywhere been noted107, that some few still existing species are common in the deposit, but have become extinct in the immediately surrounding sea; or, conversely, that some are now abundant in the neighbouring sea, but are rare or absent in this particular deposit. It is an excellent lesson to reflect on the ascertained amount of migration of the inhabitants of Europe during the Glacial period, which forms only a part of one whole geological period; and likewise to reflect on the great changes of level, on the inordinately108 great change of climate, on the prodigious lapse of time, all included within this same glacial period. Yet it may be doubted whether in any quarter of the world, sedimentary deposits, INCLUDING FOSSIL REMAINS, have gone on accumulating within the same area during the whole of this period. It is not, for instance, probable that sediment was deposited during the whole of the glacial period near the mouth of the Mississippi, within that limit of depth at which marine animals can flourish; for we know what vast geographical109 changes occurred in other parts of America during this space of time. When such beds as were deposited in shallow water near the mouth of the Mississippi during some part of the glacial period shall have been upraised, organic remains will probably first appear and disappear at different levels, owing to the migration of species and to geographical changes. And in the distant future, a geologist examining these beds, might be tempted to conclude that the average duration of life of the embedded fossils had been less than that of the glacial period, instead of having been really far greater, that is extending from before the glacial epoch110 to the present day.
In order to get a perfect gradation between two forms in the upper and lower parts of the same formation, the deposit must have gone on accumulating for a very long period, in order to have given sufficient time for the slow process of variation; hence the deposit will generally have to be a very thick one; and the species undergoing modification will have had to live on the same area throughout this whole time. But we have seen that a thick fossiliferous formation can only be accumulated during a period of subsidence; and to keep the depth approximately the same, which is necessary in order to enable the same species to live on the same space, the supply of sediment must nearly have counterbalanced the amount of subsidence. But this same movement of subsidence will often tend to sink the area whence the sediment is derived, and thus diminish the supply whilst the downward movement continues. In fact, this nearly exact balancing between the supply of sediment and the amount of subsidence is probably a rare contingency111; for it has been observed by more than one palaeontologist, that very thick deposits are usually barren of organic remains, except near their upper or lower limits.
It would seem that each separate formation, like the whole pile of formations in any country, has generally been intermittent in its accumulation. When we see, as is so often the case, a formation composed of beds of different mineralogical composition, we may reasonably suspect that the process of deposition has been much interrupted, as a change in the currents of the sea and a supply of sediment of a different nature will generally have been due to geographical changes requiring much time. Nor will the closest inspection112 of a formation give any idea of the time which its deposition has consumed. Many instances could be given of beds only a few feet in thickness, representing formations, elsewhere thousands of feet in thickness, and which must have required an enormous period for their accumulation; yet no one ignorant of this fact would have suspected the vast lapse of time represented by the thinner formation. Many cases could be given of the lower beds of a formation having been upraised, denuded, submerged, and then re-covered by the upper beds of the same formation,--facts, showing what wide, yet easily overlooked, intervals have occurred in its accumulation. In other cases we have the plainest evidence in great fossilised trees, still standing113 upright as they grew, of many long intervals of time and changes of level during the process of deposition, which would never even have been suspected, had not the trees chanced to have been preserved: thus, Messrs. Lyell and Dawson found carboniferous beds 1400 feet thick in Nova Scotia, with ancient root-bearing strata, one above the other, at no less than sixty-eight different levels. Hence, when the same species occur at the bottom, middle, and top of a formation, the probability is that they have not lived on the same spot during the whole period of deposition, but have disappeared and reappeared, perhaps many times, during the same geological period. So that if such species were to undergo a considerable amount of modification during any one geological period, a section would not probably include all the fine intermediate gradations which must on my theory have existed between them, but abrupt114, though perhaps very slight, changes of form.
It is all-important to remember that naturalists115 have no golden rule by which to distinguish species and varieties; they grant some little variability to each species, but when they meet with a somewhat greater amount of difference between any two forms, they rank both as species, unless they are enabled to connect them together by close intermediate gradations. And this from the reasons just assigned we can seldom hope to effect in any one geological section. Supposing B and C to be two species, and a third, A, to be found in an underlying bed; even if A were strictly intermediate between B and C, it would simply be ranked as a third and distinct species, unless at the same time it could be most closely connected with either one or both forms by intermediate varieties. Nor should it be forgotten, as before explained, that A might be the actual progenitor of B and C, and yet might not at all necessarily be strictly intermediate between them in all points of structure. So that we might obtain the parent-species and its several modified descendants from the lower and upper beds of a formation, and unless we obtained numerous transitional gradations, we should not recognise their relationship, and should consequently be compelled to rank them all as distinct species.
It is notorious on what excessively slight differences many palaeontologists have founded their species; and they do this the more readily if the specimens come from different sub-stages of the same formation. Some experienced conchologists are now sinking many of the very fine species of D'Orbigny and others into the rank of varieties; and on this view we do find the kind of evidence of change which on my theory we ought to find. Moreover, if we look to rather wider intervals, namely, to distinct but consecutive stages of the same great formation, we find that the embedded fossils, though almost universally ranked as specifically different, yet are far more closely allied to each other than are the species found in more widely separated formations; but to this subject I shall have to return in the following chapter.
One other consideration is worth notice: with animals and plants that can propagate rapidly and are not highly locomotive, there is reason to suspect, as we have formerly seen, that their varieties are generally at first local; and that such local varieties do not spread widely and supplant their parent-forms until they have been modified and perfected in some considerable degree. According to this view, the chance of discovering in a formation in any one country all the early stages of transition between any two forms, is small, for the successive changes are supposed to have been local or confined to some one spot. Most marine animals have a wide range; and we have seen that with plants it is those which have the widest range, that oftenest present varieties; so that with shells and other marine animals, it is probably those which have had the widest range, far exceeding the limits of the known geological formations of Europe, which have oftenest given rise, first to local varieties and ultimately to new species; and this again would greatly lessen117 the chance of our being able to trace the stages of transition in any one geological formation.
It should not be forgotten, that at the present day, with perfect specimens for examination, two forms can seldom be connected by intermediate varieties and thus proved to be the same species, until many specimens have been collected from many places; and in the case of fossil species this could rarely be effected by palaeontologists. We shall, perhaps, best perceive the improbability of our being enabled to connect species by numerous, fine, intermediate, fossil links, by asking ourselves whether, for instance, geologists at some future period will be able to prove, that our different breeds of cattle, sheep, horses, and dogs have descended from a single stock or from several aboriginal118 stocks; or, again, whether certain sea-shells inhabiting the shores of North America, which are ranked by some conchologists as distinct species from their European representatives, and by other conchologists as only varieties, are really varieties or are, as it is called, specifically distinct. This could be effected only by the future geologist discovering in a fossil state numerous intermediate gradations; and such success seems to me improbable in the highest degree.
Geological research, though it has added numerous species to existing and extinct genera, and has made the intervals between some few groups less wide than they otherwise would have been, yet has done scarcely anything in breaking down the distinction between species, by connecting them together by numerous, fine, intermediate varieties; and this not having been effected, is probably the gravest and most obvious of all the many objections which may be urged against my views. Hence it will be worth while to sum up the foregoing remarks, under an imaginary illustration. The Malay Archipelago is of about the size of Europe from the North Cape63 to the Mediterranean, and from Britain to Russia; and therefore equals all the geological formations which have been examined with any accuracy, excepting those of the United States of America. I fully agree with Mr. Godwin-Austen, that the present condition of the Malay Archipelago, with its numerous large islands separated by wide and shallow seas, probably represents the former state of Europe, when most of our formations were accumulating. The Malay Archipelago is one of the richest regions of the whole world in organic beings; yet if all the species were to be collected which have ever lived there, how imperfectly would they represent the natural history of the world!
But we have every reason to believe that the terrestrial productions of the archipelago would be preserved in an excessively imperfect manner in the formations which we suppose to be there accumulating. I suspect that not many of the strictly littoral animals, or of those which lived on naked submarine rocks, would be embedded; and those embedded in gravel or sand, would not endure to a distant epoch. Wherever sediment did not accumulate on the bed of the sea, or where it did not accumulate at a sufficient rate to protect organic bodies from decay, no remains could be preserved.
In our archipelago, I believe that fossiliferous formations could be formed of sufficient thickness to last to an age, as distant in futurity as the secondary formations lie in the past, only during periods of subsidence. These periods of subsidence would be separated from each other by enormous intervals, during which the area would be either stationary or rising; whilst rising, each fossiliferous formation would be destroyed, almost as soon as accumulated, by the incessant coast-action, as we now see on the shores of South America. During the periods of subsidence there would probably be much extinction of life; during the periods of elevation, there would be much variation, but the geological record would then be least perfect.
It may be doubted whether the duration of any one great period of subsidence over the whole or part of the archipelago, together with a contemporaneous accumulation of sediment, would EXCEED the average duration of the same specific forms; and these contingencies120 are indispensable for the preservation of all the transitional gradations between any two or more species. If such gradations were not fully preserved, transitional varieties would merely appear as so many distinct species. It is, also, probable that each great period of subsidence would be interrupted by oscillations of level, and that slight climatal changes would intervene during such lengthy121 periods; and in these cases the inhabitants of the archipelago would have to migrate, and no closely consecutive record of their modifications122 could be preserved in any one formation.
Very many of the marine inhabitants of the archipelago now range thousands of miles beyond its confines; and analogy leads me to believe that it would be chiefly these far-ranging species which would oftenest produce new varieties; and the varieties would at first generally be local or confined to one place, but if possessed of any decided123 advantage, or when further modified and improved, they would slowly spread and supplant their parent-forms. When such varieties returned to their ancient homes, as they would differ from their former state, in a nearly uniform, though perhaps extremely slight degree, they would, according to the principles followed by many palaeontologists, be ranked as new and distinct species.
If then, there be some degree of truth in these remarks, we have no right to expect to find in our geological formations, an infinite number of those fine transitional forms, which on my theory assuredly have connected all the past and present species of the same group into one long and branching chain of life. We ought only to look for a few links, some more closely, some more distantly related to each other; and these links, let them be ever so close, if found in different stages of the same formation, would, by most palaeontologists, be ranked as distinct species. But I do not pretend that I should ever have suspected how poor a record of the mutations of life, the best preserved geological section presented, had not the difficulty of our not discovering innumerable transitional links between the species which appeared at the commencement and close of each formation, pressed so hardly on my theory.
ON THE SUDDEN APPEARANCE OF WHOLE GROUPS OF ALLIED SPECIES.
The abrupt manner in which whole groups of species suddenly appear in certain formations, has been urged by several palaeontologists, for instance, by Agassiz, Pictet, and by none more forcibly than by Professor Sedgwick, as a fatal objection to the belief in the transmutation of species. If numerous species, belonging to the same genera or families, have really started into life all at once, the fact would be fatal to the theory of descent with slow modification through natural selection. For the development of a group of forms, all of which have descended from some one progenitor, must have been an extremely slow process; and the progenitors124 must have lived long ages before their modified descendants. But we continually over-rate the perfection of the geological record, and falsely infer, because certain genera or families have not been found beneath a certain stage, that they did not exist before that stage. We continually forget how large the world is, compared with the area over which our geological formations have been carefully examined; we forget that groups of species may elsewhere have long existed and have slowly multiplied before they invaded the ancient archipelagoes of Europe and of the United States. We do not make due allowance for the enormous intervals of time, which have probably elapsed between our consecutive formations,--longer perhaps in some cases than the time required for the accumulation of each formation. These intervals will have given time for the multiplication125 of species from some one or some few parent-forms; and in the succeeding formation such species will appear as if suddenly created.
I may here recall a remark formerly made, namely that it might require a long succession of ages to adapt an organism to some new and peculiar line of life, for instance to fly through the air; but that when this had been effected, and a few species had thus acquired a great advantage over other organisms, a comparatively short time would be necessary to produce many divergent forms, which would be able to spread rapidly and widely throughout the world.
I will now give a few examples to illustrate126 these remarks; and to show how liable we are to error in supposing that whole groups of species have suddenly been produced. I may recall the well-known fact that in geological treatises, published not many years ago, the great class of mammals was always spoken of as having abruptly127 come in at the commencement of the tertiary series. And now one of the richest known accumulations of fossil mammals belongs to the middle of the secondary series; and one true mammal has been discovered in the new red sandstone at nearly the commencement of this great series. Cuvier used to urge that no monkey occurred in any tertiary stratum; but now extinct species have been discovered in India, South America, and in Europe even as far back as the eocene stage. The most striking case, however, is that of the Whale family; as these animals have huge bones, are marine, and range over the world, the fact of not a single bone of a whale having been discovered in any secondary formation, seemed fully to justify128 the belief that this great and distinct order had been suddenly produced in the interval between the latest secondary and earliest tertiary formation. But now we may read in the Supplement to Lyell's 'Manual,' published in 1858, clear evidence of the existence of whales in the upper greensand, some time before the close of the secondary period.
I may give another instance, which from having passed under my own eyes has much struck me. In a memoir on Fossil Sessile Cirripedes, I have stated that, from the number of existing and extinct tertiary species; from the extraordinary abundance of the individuals of many species all over the world, from the Arctic regions to the equator, inhabiting various zones of depths from the upper tidal limits to 50 fathoms129; from the perfect manner in which specimens are preserved in the oldest tertiary beds; from the ease with which even a fragment of a valve can be recognised; from all these circumstances, I inferred that had sessile cirripedes existed during the secondary periods, they would certainly have been preserved and discovered; and as not one species had been discovered in beds of this age, I concluded that this great group had been suddenly developed at the commencement of the tertiary series. This was a sore trouble to me, adding as I thought one more instance of the abrupt appearance of a great group of species. But my work had hardly been published, when a skilful palaeontologist, M. Bosquet, sent me a drawing of a perfect specimen66 of an unmistakeable sessile cirripede, which he had himself extracted from the chalk of Belgium. And, as if to make the case as striking as possible, this sessile cirripede was a Chthamalus, a very common, large, and ubiquitous genus, of which not one specimen has as yet been found even in any tertiary stratum. Hence we now positively130 know that sessile cirripedes existed during the secondary period; and these cirripedes might have been the progenitors of our many tertiary and existing species.
The case most frequently insisted on by palaeontologists of the apparently sudden appearance of a whole group of species, is that of the teleostean fishes, low down in the Chalk period. This group includes the large majority of existing species. Lately, Professor Pictet has carried their existence one sub-stage further back; and some palaeontologists believe that certain much older fishes, of which the affinities131 are as yet imperfectly known, are really teleostean. Assuming, however, that the whole of them did appear, as Agassiz believes, at the commencement of the chalk formation, the fact would certainly be highly remarkable132; but I cannot see that it would be an insuperable difficulty on my theory, unless it could likewise be shown that the species of this group appeared suddenly and simultaneously133 throughout the world at this same period. It is almost superfluous to remark that hardly any fossil-fish are known from south of the equator; and by running through Pictet's Palaeontology it will be seen that very few species are known from several formations in Europe. Some few families of fish now have a confined range; the teleostean fish might formerly have had a similarly confined range, and after having been largely developed in some one sea, might have spread widely. Nor have we any right to suppose that the seas of the world have always been so freely open from south to north as they are at present. Even at this day, if the Malay Archipelago were converted into land, the tropical parts of the Indian Ocean would form a large and perfectly119 enclosed basin, in which any great group of marine animals might be multiplied; and here they would remain confined, until some of the species became adapted to a cooler climate, and were enabled to double the southern capes134 of Africa or Australia, and thus reach other and distant seas.
From these and similar considerations, but chiefly from our ignorance of the geology of other countries beyond the confines of Europe and the United States; and from the revolution in our palaeontological ideas on many points, which the discoveries of even the last dozen years have effected, it seems to me to be about as rash in us to dogmatize on the succession of organic beings throughout the world, as it would be for a naturalist116 to land for five minutes on some one barren point in Australia, and then to discuss the number and range of its productions.
ON THE SUDDEN APPEARANCE OF GROUPS OF ALLIED SPECIES IN THE LOWEST KNOWN FOSSILIFEROUS STRATA.
There is another and allied difficulty, which is much graver. I allude135 to the manner in which numbers of species of the same group, suddenly appear in the lowest known fossiliferous rocks. Most of the arguments which have convinced me that all the existing species of the same group have descended from one progenitor, apply with nearly equal force to the earliest known species. For instance, I cannot doubt that all the Silurian trilobites have descended from some one crustacean136, which must have lived long before the Silurian age, and which probably differed greatly from any known animal. Some of the most ancient Silurian animals, as the Nautilus, Lingula, etc., do not differ much from living species; and it cannot on my theory be supposed, that these old species were the progenitors of all the species of the orders to which they belong, for they do not present characters in any degree intermediate between them. If, moreover, they had been the progenitors of these orders, they would almost certainly have been long ago supplanted137 and exterminated by their numerous and improved descendants.
Consequently, if my theory be true, it is indisputable that before the lowest Silurian stratum was deposited, long periods elapsed, as long as, or probably far longer than, the whole interval from the Silurian age to the present day; and that during these vast, yet quite unknown, periods of time, the world swarmed138 with living creatures.
To the question why we do not find records of these vast primordial139 periods, I can give no satisfactory answer. Several of the most eminent140 geologists, with Sir R. Murchison at their head, are convinced that we see in the organic remains of the lowest Silurian stratum the dawn of life on this planet. Other highly competent judges, as Lyell and the late E. Forbes, dispute this conclusion. We should not forget that only a small portion of the world is known with accuracy. M. Barrande has lately added another and lower stage to the Silurian system, abounding141 with new and peculiar species. Traces of life have been detected in the Longmynd beds beneath Barrande's so-called primordial zone. The presence of phosphatic nodules and bituminous matter in some of the lowest azoic rocks, probably indicates the former existence of life at these periods. But the difficulty of understanding the absence of vast piles of fossiliferous strata, which on my theory no doubt were somewhere accumulated before the Silurian epoch, is very great. If these most ancient beds had been wholly worn away by denudation, or obliterated142 by metamorphic action, we ought to find only small remnants of the formations next succeeding them in age, and these ought to be very generally in a metamorphosed condition. But the descriptions which we now possess of the Silurian deposits over immense territories in Russia and in North America, do not support the view, that the older a formation is, the more it has suffered the extremity of denudation and metamorphism.
The case at present must remain inexplicable143; and may be truly urged as a valid144 argument against the views here entertained. To show that it may hereafter receive some explanation, I will give the following hypothesis. From the nature of the organic remains, which do not appear to have inhabited profound depths, in the several formations of Europe and of the United States; and from the amount of sediment, miles in thickness, of which the formations are composed, we may infer that from first to last large islands or tracts145 of land, whence the sediment was derived, occurred in the neighbourhood of the existing continents of Europe and North America. But we do not know what was the state of things in the intervals between the successive formations; whether Europe and the United States during these intervals existed as dry land, or as a submarine surface near land, on which sediment was not deposited, or again as the bed of an open and unfathomable sea.
Looking to the existing oceans, which are thrice as extensive as the land, we see them studded with many islands; but not one oceanic island is as yet known to afford even a remnant of any palaeozoic or secondary formation. Hence we may perhaps infer, that during the palaeozoic and secondary periods, neither continents nor continental146 islands existed where our oceans now extend; for had they existed there, palaeozoic and secondary formations would in all probability have been accumulated from sediment derived from their wear and tear; and would have been at least partially upheaved by the oscillations of level, which we may fairly conclude must have intervened during these enormously long periods. If then we may infer anything from these facts, we may infer that where our oceans now extend, oceans have extended from the remotest period of which we have any record; and on the other hand, that where continents now exist, large tracts of land have existed, subjected no doubt to great oscillations of level, since the earliest silurian period. The coloured map appended to my volume on Coral Reefs, led me to conclude that the great oceans are still mainly areas of subsidence, the great archipelagoes still areas of oscillations of level, and the continents areas of elevation. But have we any right to assume that things have thus remained from eternity? Our continents seem to have been formed by a preponderance, during many oscillations of level, of the force of elevation; but may not the areas of preponderant movement have changed in the lapse of ages? At a period immeasurably antecedent to the silurian epoch, continents may have existed where oceans are now spread out; and clear and open oceans may have existed where our continents now stand. Nor should we be justified147 in assuming that if, for instance, the bed of the Pacific Ocean were now converted into a continent, we should there find formations older than the silurian strata, supposing such to have been formerly deposited; for it might well happen that strata which had subsided148 some miles nearer to the centre of the earth, and which had been pressed on by an enormous weight of superincumbent water, might have undergone far more metamorphic action than strata which have always remained nearer to the surface. The immense areas in some parts of the world, for instance in South America, of bare metamorphic rocks, which must have been heated under great pressure, have always seemed to me to require some special explanation; and we may perhaps believe that we see in these large areas, the many formations long anterior149 to the silurian epoch in a completely metamorphosed condition.
The several difficulties here discussed, namely our not finding in the successive formations infinitely numerous transitional links between the many species which now exist or have existed; the sudden manner in which whole groups of species appear in our European formations; the almost entire absence, as at present known, of fossiliferous formations beneath the Silurian strata, are all undoubtedly150 of the gravest nature. We see this in the plainest manner by the fact that all the most eminent palaeontologists, namely Cuvier, Owen, Agassiz, Barrande, Falconer, E. Forbes, etc., and all our greatest geologists, as Lyell, Murchison, Sedgwick, etc., have unanimously, often vehemently151, maintained the immutability152 of species. But I have reason to believe that one great authority, Sir Charles Lyell, from further reflexion entertains grave doubts on this subject. I feel how rash it is to differ from these great authorities, to whom, with others, we owe all our knowledge. Those who think the natural geological record in any degree perfect, and who do not attach much weight to the facts and arguments of other kinds given in this volume, will undoubtedly at once reject my theory. For my part, following out Lyell's metaphor153, I look at the natural geological record, as a history of the world imperfectly kept, and written in a changing dialect; of this history we possess the last volume alone, relating only to two or three countries. Of this volume, only here and there a short chapter has been preserved; and of each page, only here and there a few lines. Each word of the slowly-changing language, in which the history is supposed to be written, being more or less different in the interrupted succession of chapters, may represent the apparently abruptly changed forms of life, entombed in our consecutive, but widely separated formations. On this view, the difficulties above discussed are greatly diminished, or even disappear.
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1
lapse
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| n.过失，流逝，失效，抛弃信仰，间隔；vi.堕落，停止，失效，流逝；vt.使失效 | |
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deposition
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| n.免职，罢官；作证；沉淀；沉淀物 | |
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denudation
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| n.剥下；裸露；滥伐；剥蚀 | |
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intermittence
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| n.间断；间歇 | |
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strata
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| n.地层（复数）；社会阶层 | |
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enumerated
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| v.列举，枚举，数( enumerate的过去式和过去分词 ) | |
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apparently
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| adv.显然地；表面上，似乎 | |
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favourable
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| adj.赞成的，称赞的，有利的，良好的，顺利的 | |
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lesser
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| adj.次要的，较小的；adv.较小地，较少地 | |
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exterminated
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| v.消灭，根绝( exterminate的过去式和过去分词 ) | |
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exterminate
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| v.扑灭，消灭，根绝 | |
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modification
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| n.修改，改进，缓和，减轻 | |
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extermination
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| n.消灭，根绝 | |
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formerly
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| adv.从前，以前 | |
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stratum
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| n.地层，社会阶层 | |
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progenitor
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| n.祖先，先驱 | |
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descended
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| a.为...后裔的，出身于...的 | |
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possessed
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| adj.疯狂的；拥有的，占有的 | |
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determined
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| adj.坚定的；有决心的 | |
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mere
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| adj.纯粹的；仅仅，只不过 | |
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allied
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| adj.协约国的；同盟国的 | |
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organisation
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| n.组织，安排，团体，有机休 | |
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considerably
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| adv.极大地；相当大地；在很大程度上 | |
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supplant
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| vt.排挤；取代 | |
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backwards
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| adv.往回地，向原处，倒，相反，前后倒置地 | |
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converging
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| adj.收敛[缩]的，会聚的，趋同的v.（线条、运动的物体等）会于一点( converge的现在分词 )；（趋于）相似或相同；人或车辆汇集；聚集 | |
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remains
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| n.剩余物，残留物；遗体，遗迹 | |
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infinitely
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| adv.无限地，无穷地 | |
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geologist
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| n.地质学家 | |
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treatises
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| n.专题著作，专题论文，专著( treatise的名词复数 ) | |
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inadequate
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| adj.(for,to)不充足的，不适当的 | |
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sediment
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| n.沉淀，沉渣，沉积(物) | |
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degradation
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| n.降级；低落；退化；陵削；降解；衰变 | |
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pebbles
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| [复数]鹅卵石； 沙砾； 卵石，小圆石( pebble的名词复数 ) | |
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fixed
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| adj.固定的，不变的，准备好的；(计算机)固定的 | |
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boulders
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| n.卵石( boulder的名词复数 )；巨砾；（受水或天气侵蚀而成的）巨石；漂砾 | |
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marine
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| adj.海的；海生的；航海的；海事的；n.水兵 | |
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abraded
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| adj.[医]刮擦的v.刮擦( abrade的过去式和过去分词 )；（在精神方面）折磨（人）；消磨（意志、精神等）；使精疲力尽 | |
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promontory
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| n.海角；岬 | |
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miller
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| n.磨坊主 | |
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conglomerate
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| n.综合商社，多元化集团公司 | |
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igneous
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| adj.火的，火绒的 | |
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geologists
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| 地质学家，地质学者( geologist的名词复数 ) | |
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volcanic
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| adj.火山的；象火山的；由火山引起的 | |
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perpendicular
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| adj.垂直的，直立的；n.垂直线，垂直的位置 | |
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upwards
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| adv.向上，在更高处...以上 | |
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vertical
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| adj.垂直的，顶点的，纵向的；n.垂直物，垂直的位置 | |
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displacement
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| n.移置，取代，位移，排水量 | |
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varied
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| adj.多样的，多变化的 | |
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fully
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| adv.完全地，全部地，彻底地；充分地 | |
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prodigious
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| adj.惊人的，奇妙的；异常的；巨大的；庞大的 | |
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smoothly
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| adv.平滑地，顺利地，流利地，流畅地 | |
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eternity
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| n.不朽，来世；永恒，无穷 | |
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tempted
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| v.怂恿（某人）干不正当的事；冒…的险（tempt的过去分词） | |
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memoir
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| n.[pl.]回忆录，自传；记事录 | |
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dome
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| n.圆屋顶，拱顶 | |
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underlies
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| v.位于或存在于（某物）之下( underlie的第三人称单数 )；构成…的基础（或起因），引起 | |
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applied
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| adj.应用的；v.应用，适用 | |
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extremity
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| n.末端，尽头；尽力；终极；极度 | |
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requisite
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| adj.需要的，必不可少的；n.必需品 | |
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61
denuded
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| adj.[医]变光的，裸露的v.使赤裸( denude的过去式和过去分词 )；剥光覆盖物 | |
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indented
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| adj.锯齿状的，高低不平的；缩进排版 | |
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cape
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| n.海角，岬；披肩，短披风 | |
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paltry
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| adj.无价值的，微不足道的 | |
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behold
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| v.看，注视，看到 | |
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specimen
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| n.样本，标本 | |
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specimens
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| n.样品( specimen的名词复数 )；范例；（化验的）抽样；某种类型的人 | |
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sufficiently
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| adv.足够地，充分地 | |
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embed
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| vt.把…嵌(埋、插)入，扎牢；使深留脑中 | |
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tint
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| n.淡色，浅色；染发剂；vt.着以淡淡的颜色 | |
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bespeaks
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| v.预定( bespeak的第三人称单数 )；订（货）；证明；预先请求 | |
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interval
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| n.间隔，间距；幕间休息，中场休息 | |
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underlying
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| adj.在下面的，含蓄的，潜在的 | |
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embedded
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| a.扎牢的 | |
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gravel
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| n.砂跞；砂砾层；结石 | |
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percolation
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| n.过滤，浸透；渗滤；渗漏 | |
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strictly
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| adv.严厉地，严格地；严密地 | |
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littoral
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| adj.海岸的；湖岸的；n.沿（海）岸地区 | |
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Mediterranean
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| adj.地中海的；地中海沿岸的 | |
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partially
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| adv.部分地，从某些方面讲 | |
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analogous
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| adj.相似的；类似的 | |
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82
superfluous
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| adj.过多的，过剩的，多余的 | |
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preservation
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| n.保护，维护，保存，保留，保持 | |
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intervals
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| n.[军事]间隔( interval的名词复数 )；间隔时间；[数学]区间；（戏剧、电影或音乐会的）幕间休息 | |
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85
tabulated
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| 把（数字、事实）列成表( tabulate的过去式和过去分词 ) | |
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consecutive
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| adj.连续的，联贯的，始终一贯的 | |
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skilful
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| (=skillful)adj.灵巧的，熟练的 | |
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peculiar
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| adj.古怪的，异常的；特殊的，特有的 | |
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ascertained
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| v.弄清，确定，查明( ascertain的过去式和过去分词 ) | |
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derived
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| vi.起源；由来；衍生；导出v.得到( derive的过去式和过去分词 )；（从…中）得到获得；源于；（从…中）提取 | |
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intermittent
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| adj.间歇的，断断续续的 | |
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fauna
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| n.(一个地区或时代的)所有动物，动物区系 | |
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scantily
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| adv.缺乏地；不充足地；吝啬地；狭窄地 | |
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faunas
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| 动物群 | |
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incessant
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| adj.不停的，连续的 | |
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subside
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| vi.平静，平息；下沉，塌陷，沉降 | |
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affected
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| adj.不自然的，假装的 | |
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stationary
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| adj.固定的，静止不动的 | |
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elevation
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| n.高度；海拔；高地；上升；提高 | |
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accurately
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| adv.准确地，精确地 | |
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previously
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| adv.以前，先前（地） | |
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extinction
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| n.熄灭，消亡，消灭，灭绝，绝种 | |
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worthy
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| adj.(of)值得的，配得上的；有价值的 | |
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deference
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| n.尊重，顺从；敬意 | |
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migration
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| n.迁移，移居，（鸟类等的）迁徙 | |
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immigrated
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| v.移入( immigrate的过去式和过去分词 )；移民 | |
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noted
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| adj.著名的，知名的 | |
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inordinately
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| adv.无度地，非常地 | |
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geographical
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| adj.地理的；地区(性)的 | |
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epoch
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| n.(新)时代；历元 | |
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111
contingency
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| n.意外事件，可能性 | |
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112
inspection
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| n.检查，审查，检阅 | |
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113
standing
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| n.持续，地位；adj.永久的，不动的，直立的，不流动的 | |
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abrupt
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| adj.突然的，意外的；唐突的，鲁莽的 | |
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115
naturalists
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| n.博物学家( naturalist的名词复数 )；（文学艺术的）自然主义者 | |
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116
naturalist
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| n.博物学家(尤指直接观察动植物者) | |
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117
lessen
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| vt.减少，减轻；缩小 | |
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118
aboriginal
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| adj.（指动植物）土生的，原产地的，土著的 | |
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perfectly
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| adv.完美地，无可非议地，彻底地 | |
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contingencies
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| n.偶然发生的事故，意外事故( contingency的名词复数 )；以备万一 | |
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121
lengthy
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| adj.漫长的，冗长的 | |
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122
modifications
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| n.缓和( modification的名词复数 )；限制；更改；改变 | |
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123
decided
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| adj.决定了的，坚决的；明显的，明确的 | |
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progenitors
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| n.祖先( progenitor的名词复数 )；先驱；前辈；原本 | |
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multiplication
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| n.增加，增多，倍增；增殖，繁殖；乘法 | |
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126
illustrate
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| v.举例说明，阐明；图解，加插图 | |
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127
abruptly
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| adv.突然地，出其不意地 | |
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128
justify
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| vt.证明…正当(或有理)，为…辩护 | |
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129
fathoms
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| 英寻( fathom的名词复数 ) | |
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130
positively
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| adv.明确地，断然，坚决地；实在，确实 | |
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131
affinities
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| n.密切关系( affinity的名词复数 )；亲近；（生性）喜爱；类同 | |
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132
remarkable
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| adj.显著的，异常的，非凡的，值得注意的 | |
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133
simultaneously
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| adv.同时发生地，同时进行地 | |
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134
capes
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| 碎谷； 斗篷( cape的名词复数 )； 披肩； 海角； 岬 | |
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135
allude
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| v.提及，暗指 | |
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136
crustacean
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| n.甲壳动物；adj.甲壳纲的 | |
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137
supplanted
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| 把…排挤掉，取代( supplant的过去式和过去分词 ) | |
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138
swarmed
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| 密集( swarm的过去式和过去分词 )； 云集； 成群地移动； 蜜蜂或其他飞行昆虫成群地飞来飞去 | |
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139
primordial
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| adj.原始的；最初的 | |
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140
eminent
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| adj.显赫的，杰出的，有名的，优良的 | |
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141
abounding
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| adj.丰富的，大量的v.大量存在，充满，富于( abound的现在分词 ) | |
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142
obliterated
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| v.除去( obliterate的过去式和过去分词 )；涂去；擦掉；彻底破坏或毁灭 | |
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143
inexplicable
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| adj.无法解释的，难理解的 | |
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144
valid
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| adj.有确实根据的；有效的；正当的，合法的 | |
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145
tracts
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| 大片土地( tract的名词复数 )； 地带； （体内的）道； （尤指宣扬宗教、伦理或政治的）短文 | |
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146
continental
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| adj.大陆的，大陆性的，欧洲大陆的 | |
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147
justified
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| a.正当的，有理的 | |
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148
subsided
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| v.（土地）下陷（因在地下采矿）( subside的过去式和过去分词 )；减弱；下降至较低或正常水平；一下子坐在椅子等上 | |
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149
anterior
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| adj.较早的；在前的 | |
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150
undoubtedly
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| adv.确实地，无疑地 | |
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151
vehemently
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| adv. 热烈地 | |
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152
immutability
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| n.不变(性) | |
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153
metaphor
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| n.隐喻，暗喻 | |
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