Chapter II

On the Motions of the Heart as Seen in the Dissection¹ of Living Animals

In the first place, then, when the chest of a living animal is laid open and the capsule that immediately surrounds the heart is slit² up or removed, the organ is seen now to move, now to be at rest; there is a time when it moves, and a time when it is motionless.

These things are more obvious in the colder animals, such as toads³, frogs, serpents, small fishes, crabs⁴, shrimps⁵, snails⁶, and shell-fish. They also become more distinct in warm-blooded animals, such as the dog and hog⁷, if they be attentively⁸ noted⁹ when the heart begins to flag, to move more slowly, and, as it were, to die: the movements then become slower and rarer, the pauses longer, by which it is made much more easy to perceive and unravel¹⁰ what the motions really are, and how they are performed. In the pause, as in death, the heart is soft, flaccid, exhausted¹¹, lying, as it were, at rest.

In the motion, and interval¹² in which this is accomplished¹³, three principal circumstances are to be noted:

1. That the heart is erected¹⁵, and rises upwards¹⁶ to a point, so that at this time it strikes against the breast and the pulse is felt externally.

2. That it is everywhere contracted, but more especially towards the sides so that it looks narrower, relatively¹⁷ longer, more drawn¹⁸ together. The heart of an eel¹⁹ taken out of the body of the animal and placed upon the table or the hand, shows these particulars; but the same things are manifest in the hearts of all small fishes and of those colder animals where the organ is more conical or elongated²⁰.

3. The heart being grasped in the hand, is felt to become harder during its action. Now this hardness proceeds from tension, precisely²¹ as when the forearm is grasped, its tendons are perceived to become tense and resilient when the fingers are moved.

4. It may further be observed in fishes, and the colder blooded animals, such as frogs, serpents, etc., that the heart, when it moves, becomes of a paler color, when quiescent²² of a deeper blood-red color.

From these particulars it appears evident to me that the motion of the heart consists in a certain universal tension — both contraction²³ in the line of its fibres, and constriction²⁴ in every sense. It becomes erect¹⁴, hard, and of diminished size during its action; the motion is plainly of the same nature as that of the muscles when they contract in the line of their sinews and fibres; for the muscles, when in action, acquire vigor²⁵ and tenseness, and from soft become hard, prominent, and thickened: and in the same manner the heart.

We are therefore authorized²⁶ to conclude that the heart, at the moment of its action, is at once constricted²⁷ on all sides, rendered thicker in its parietes and smaller in its ventricles, and so made apt to project or expel its charge of blood. This, indeed, is made sufficiently²⁸ manifest by the preceding fourth observation in which we have seen that the heart, by squeezing out the blood that it contains, becomes paler, and then when it sinks into repose²⁹ and the ventricle is filled anew with blood, that the deeper crimson³⁰ colour returns. But no one need remain in doubt of the fact, for if the ventricle be pierced the blood will be seen to be forcibly projected outwards³¹ upon each motion or pulsation³² when the heart is tense.

These things, therefore, happen together or at the same instant: the tension of the heart, the pulse of its apex³³, which is felt externally by its striking against the chest, the thickening of its parietes, and the forcible expulsion of the blood it contains by the constriction of its ventricles.

Hence the very opposite of the opinions commonly received appears to be true; inasmuch as it is generally believed that when the heart strikes the breast and the pulse is felt without, the heart is dilated³⁴ in its ventricles and is filled with blood; but the contrary of this is the fact, and the heart, when it contracts (and the impulse of the apex is conveyed through the chest wall), is emptied. Whence the motion which is generally regarded as the diastole of the heart, is in truth its systole. And in like manner the intrinsic motion of the heart is not the diastole but the systole; neither is it in the diastole that the heart grows firm and tense, but in the systole, for then only, when tense, is it moved and made vigorous.

Neither is it by any means to be allowed that the heart only moves in the lines of its straight fibres, although the great Vesalius giving this notion countenance³⁶, quotes a bundle of osiers bound in a pyramidal heap in illustration; meaning, that as the apex is approached to the base, so are the sides made to bulge³⁷ out in the fashion of arches, the cavities to dilate³⁵, the ventricles to acquire the form of a cupping-glass and so to suck in the blood. But the true effect of every one of its fibres is to constringe the heart at the same time they render it tense; and this rather with the effect of thickening and amplifying³⁸ the walls and substance of the organ than enlarging its ventricles. And, again, as the fibres run from the apex to the base, and draw the apex towards the base, they do not tend to make the walls of the heart bulge out in circles, but rather the contrary; inasmuch as every fibre that is circularly disposed, tends to become straight when it contracts; and is distended³⁹ laterally⁴⁰ and thickened, as in the case of muscular fibres in general, when they contract, that is, when they are shortened longitudinally, as we see them in the bellies⁴¹ of the muscles of the body at large. To all this let it be added, that not only are the ventricles contracted in virtue⁴² of the direction and condensation⁴³ of their walls, but farther, that those fibres, or bands, styled nerves by Aristotle, which are so conspicuous⁴⁴ in the ventricles of the larger animals, and contain all the straight fibres (the parietes of the heart containing only circular ones), when they contract simultaneously⁴⁵ by an admirable adjustment all the internal surfaces are drawn together as if with cords, and so is the charge of blood expelled with force.

Neither is it true, as vulgarly believed, that the heart by any dilatation or motion of its own, has the power of drawing the blood into the ventricles; for when it acts and becomes tense, the blood is expelled; when it relaxes and sinks together it receives the blood in the manner and wise which will by-and-by be explained.

1 dissection   

n.分析；解剖

参考例句：

• A dissection of your argument shows several inconsistencies.对你论点作仔细分析后发现一些前后矛盾之处。
• Researchers need a growing supply of corpses for dissection.研究人员需要更多的供解剖用的尸体。

2 slit   

n.狭长的切口；裂缝；vt.切开，撕裂

参考例句：

• The coat has been slit in two places.这件外衣有两处裂开了。
• He began to slit open each envelope.他开始裁开每个信封。

3 toads   

n.蟾蜍，癞蛤蟆( toad的名词复数 )

参考例句：

• All toads blink when they swallow. 所有的癞蛤蟆吞食东西时都会眨眼皮。 来自《简明英汉词典》
• Toads have shorter legs and are generally more clumsy than frogs. 蟾蜍比青蛙脚短，一般说来没有青蛙灵活。 来自辞典例句

4 crabs   

n.蟹( crab的名词复数 )；阴虱寄生病；蟹肉v.捕蟹( crab的第三人称单数 )

参考例句：

• As we walked along the seashore we saw lots of tiny crabs. 我们在海岸上散步时看到很多小蟹。 来自《简明英汉词典》
• The fish and crabs scavenge for decaying tissue. 鱼和蟹搜寻腐烂的组织为食。 来自《简明英汉词典》

5 shrimps   

n.虾，小虾( shrimp的名词复数 )；矮小的人

参考例句：

• Shrimps are a popular type of seafood. 小虾是比较普遍的一种海味。 来自《简明英汉词典》
• I'm going to have shrimps for my tea. 傍晚的便餐我要吃点虾。 来自辞典例句

6 snails   

n.蜗牛；迟钝的人；蜗牛( snail的名词复数 )

参考例句：

• I think I'll try the snails for lunch—I'm feeling adventurous today. 我想我午餐要尝一下蜗牛——我今天很想冒险。 来自《简明英汉词典》
• Most snails have shells on their backs. 大多数蜗牛背上有壳。 来自《简明英汉词典》

7 hog   

n.猪；馋嘴贪吃的人；vt.把…占为己有，独占

参考例句：

• He is greedy like a hog.他像猪一样贪婪。
• Drivers who hog the road leave no room for other cars.那些占着路面的驾驶员一点余地都不留给其他车辆。

8 attentively   

adv.聚精会神地；周到地；谛；凝神

参考例句：

• She listened attentively while I poured out my problems. 我倾吐心中的烦恼时，她一直在注意听。 来自《简明英汉词典》
• She listened attentively and set down every word he said. 她专心听着，把他说的话一字不漏地记下来。 来自《简明英汉词典》

9 noted   

adj.著名的，知名的

参考例句：

• The local hotel is noted for its good table.当地的那家酒店以餐食精美而著称。
• Jim is noted for arriving late for work.吉姆上班迟到出了名。

10 unravel   

v.弄清楚（秘密）；拆开，解开，松开

参考例句：

• He was good with his hands and could unravel a knot or untangle yarn that others wouldn't even attempt.他的手很灵巧，其他人甚至都不敢尝试的一些难解的绳结或缠在一起的纱线，他都能解开。
• This is the attitude that led him to unravel a mystery that long puzzled Chinese historians.正是这种态度使他解决了长期以来使中国历史学家们大惑不解的谜。

11 exhausted   

adj.极其疲惫的，精疲力尽的

参考例句：

• It was a long haul home and we arrived exhausted.搬运回家的这段路程特别长，到家时我们已筋疲力尽。
• Jenny was exhausted by the hustle of city life.珍妮被城市生活的忙乱弄得筋疲力尽。

12 interval   

n.间隔，间距；幕间休息，中场休息

参考例句：

• The interval between the two trees measures 40 feet.这两棵树的间隔是40英尺。
• There was a long interval before he anwsered the telephone.隔了好久他才回了电话。

13 accomplished   

adj.有才艺的；有造诣的；达到了的

参考例句：

• Thanks to your help,we accomplished the task ahead of schedule.亏得你们帮忙,我们才提前完成了任务。
• Removal of excess heat is accomplished by means of a radiator.通过散热器完成多余热量的排出。

14 erect   

n./v.树立，建立，使竖立；adj.直立的，垂直的

参考例句：

• She held her head erect and her back straight.她昂着头，把背挺得笔直。
• Soldiers are trained to stand erect.士兵们训练站得笔直。

15 ERECTED   

adj. 直立的，竖立的，笔直的 vt. 使 ... 直立，建立

参考例句：

• A monument to him was erected in St Paul's Cathedral. 在圣保罗大教堂为他修了一座纪念碑。
• A monument was erected to the memory of that great scientist. 树立了一块纪念碑纪念那位伟大的科学家。

16 upwards   

adv.向上，在更高处...以上

参考例句：

• The trend of prices is still upwards.物价的趋向是仍在上涨。
• The smoke rose straight upwards.烟一直向上升。

17 relatively   

adv.比较...地，相对地

参考例句：

• The rabbit is a relatively recent introduction in Australia.兔子是相对较新引入澳大利亚的物种。
• The operation was relatively painless.手术相对来说不痛。

18 drawn   

v.拖，拉，拔出；adj.憔悴的，紧张的

参考例句：

• All the characters in the story are drawn from life.故事中的所有人物都取材于生活。
• Her gaze was drawn irresistibly to the scene outside.她的目光禁不住被外面的风景所吸引。

19 eel   

n.鳗鲡

参考例句：

• He used an eel spear to catch an eel.他用一只捕鳗叉捕鳗鱼。
• In Suzhou,there was a restaurant that specialized in eel noodles.苏州有一家饭馆，他们那里的招牌菜是鳗鱼面。

20 elongated   

v.延长，加长( elongate的过去式和过去分词 )

参考例句：

• Modigliani's women have strangely elongated faces. 莫迪里阿尼画中的妇女都长着奇长无比的脸。
• A piece of rubber can be elongated by streching. 一块橡皮可以拉长。 来自《用法词典》

21 precisely   

adv.恰好，正好，精确地，细致地

参考例句：

• It's precisely that sort of slick sales-talk that I mistrust.我不相信的正是那种油腔滑调的推销宣传。
• The man adjusted very precisely.那个人调得很准。

22 quiescent   

adj.静止的，不活动的，寂静的

参考例句：

• It is unlikely that such an extremist organization will remain quiescent for long.这种过激的组织是不太可能长期沉默的。
• Great distance in either time or space has wonderful power to lull and render quiescent the human mind.时间和空间上的远距离有一种奇妙的力量，可以使人的心灵平静。

23 contraction   

n.缩略词，缩写式，害病

参考例句：

• The contraction of this muscle raises the lower arm.肌肉的收缩使前臂抬起。
• The forces of expansion are balanced by forces of contraction.扩张力和收缩力相互平衡。

24 constriction   

压缩； 紧压的感觉； 束紧； 压缩物

参考例句：

• She feels a constriction in the chest. 她胸部有压迫感。
• If you strain to run fast, you start coughing and feel a constriction in the chest. 还是别跑紧了，一咬牙就咳嗽，心口窝辣蒿蒿的！ 来自汉英文学 - 骆驼祥子

25 vigor   

n.活力，精力，元气

参考例句：

• The choir sang the words out with great vigor.合唱团以极大的热情唱出了歌词。
• She didn't want to be reminded of her beauty or her former vigor.现在，她不愿人们提起她昔日的美丽和以前的精力充沛。

26 authorized   

a.委任的，许可的

参考例句：

• An administrative order is valid if authorized by a statute.如果一个行政命令得到一个法规的认可那么这个命令就是有效的。

27 constricted   

adj.抑制的，约束的

参考例句：

• Her throat constricted and she swallowed hard. 她喉咙发紧，使劲地咽了一下唾沫。
• The tight collar constricted his neck. 紧领子勒着他的脖子。

28 sufficiently   

adv.足够地，充分地

参考例句：

• It turned out he had not insured the house sufficiently.原来他没有给房屋投足保险。
• The new policy was sufficiently elastic to accommodate both views.新政策充分灵活地适用两种观点。

29 repose   

v.(使)休息；n.安息

参考例句：

• Don't disturb her repose.不要打扰她休息。
• Her mouth seemed always to be smiling,even in repose.她的嘴角似乎总是挂着微笑，即使在睡眠时也是这样。

30 crimson   

n./adj.深(绯)红色(的)；vi.脸变绯红色

参考例句：

• She went crimson with embarrassment.她羞得满脸通红。
• Maple leaves have turned crimson.枫叶已经红了。

31 outwards   

adj.外面的，公开的，向外的；adv.向外；n.外形

参考例句：

• Does this door open inwards or outwards?这门朝里开还是朝外开？
• In lapping up a fur,they always put the inner side outwards.卷毛皮时，他们总是让内层朝外。

32 pulsation   

n.脉搏，悸动，脉动；搏动性

参考例句：

• At low frequencies, such as 10 per sec., pulsation is sensed rather than vibration. 在低频率（譬如每秒十次）时，所感觉到的是脉冲而非振动。 来自辞典例句
• If the roller pulsation, the pressure on paper as cause misregister. 如果滚子径向跳不静，则差纸的不张辛有不小有小，致使套印禁绝。 来自互联网

33 apex   

n.顶点，最高点

参考例句：

• He reached the apex of power in the early 1930s.他在三十年代初达到了权力的顶峰。
• His election to the presidency was the apex of his career.当选总统是他一生事业的顶峰。

34 dilated   

adj.加宽的，扩大的v.（使某物）扩大，膨胀，张大( dilate的过去式和过去分词 )

参考例句：

• Her eyes dilated with fear. 她吓得瞪大了眼睛。
• The cat dilated its eyes. 猫瞪大了双眼。 来自《简明英汉词典》

35 dilate   

vt.使膨胀，使扩大

参考例句：

• At night,the pupils dilate to allow in more light.到了晚上，瞳孔就会扩大以接收更多光线。
• Exercise dilates blood vessels on the surface of the brain.运动会使大脑表层的血管扩张。

36 countenance   

n.脸色，面容；面部表情；vt.支持，赞同

参考例句：

• At the sight of this photograph he changed his countenance.他一看见这张照片脸色就变了。
• I made a fierce countenance as if I would eat him alive.我脸色恶狠狠地，仿佛要把他活生生地吞下去。

37 bulge   

n.突出，膨胀，激增；vt.突出，膨胀

参考例句：

• The apple made a bulge in his pocket.苹果把他口袋塞得鼓了起来。
• What's that awkward bulge in your pocket?你口袋里那块鼓鼓囊囊的东西是什么？

38 amplifying   

放大，扩大( amplify的现在分词 )； 增强； 详述

参考例句：

• Often they use borrowed funds, amplifying their gains and losses. 他们通常会用借贷的资金交易，从而放大收益或损失。
• An amplifying type (or analog) device, as opposed to digital device. 放大器类（或模拟）器件，相对于数字器件而言的。

39 distended   

v.（使）膨胀，肿胀( distend的过去式和过去分词 )

参考例句：

• starving children with huge distended bellies 鼓着浮肿肚子的挨饿儿童
• The balloon was distended. 气球已膨胀。 来自《现代英汉综合大词典》

40 laterally   

ad.横向地；侧面地；旁边地

参考例句：

• Shafts were sunk, with tunnels dug laterally. 竖井已经打下，并且挖有横向矿道。
• When the plate becomes unstable, it buckles laterally. 当板失去稳定时，就发生横向屈曲。

41 bellies   

n.肚子( belly的名词复数 )；腹部；（物体的）圆形或凸起部份；腹部…形的

参考例句：

• They crawled along on their bellies. 他们匍匐前进。
• starving children with huge distended bellies 鼓着浮肿肚子的挨饿儿童

42 virtue   

n.德行，美德；贞操；优点；功效，效力

参考例句：

• He was considered to be a paragon of virtue.他被认为是品德尽善尽美的典范。
• You need to decorate your mind with virtue.你应该用德行美化心灵。

43 condensation   

n.压缩，浓缩；凝结的水珠

参考例句：

• A cloud is a condensation of water vapour in the atmosphere.云是由大气中的水蒸气凝结成的。
• He used his sleeve to wipe the condensation off the glass.他用袖子擦掉玻璃上凝结的水珠。

44 conspicuous   

adj.明眼的，惹人注目的；炫耀的，摆阔气的

参考例句：

• It is conspicuous that smoking is harmful to health.很明显，抽烟对健康有害。
• Its colouring makes it highly conspicuous.它的色彩使它非常惹人注目。

45 simultaneously   

adv.同时发生地，同时进行地

参考例句：

• The radar beam can track a number of targets almost simultaneously.雷达波几乎可以同时追着多个目标。
• The Windows allow a computer user to execute multiple programs simultaneously.Windows允许计算机用户同时运行多个程序。